Eggs (Genus: 81; Species: 185)
VETEROVATA
(Parataxonomy of fossil eggs and shells, nests?)
NESTS
oofamily: Indet.
Oogenus: Oolithus (Oolithes) non BUCKMAN, 1859
O. sphaericus van STRAELEN, 1928
O. megadermus YOUNG, 1959
O. nanhsiungensis YOUNG, 1965
Oogenus: Parvoblongoolithus ZHANG, JIN, O’CONNOR, WANG & XIE, 2014
P. jinguoensis ZHANG, JIN, O’CONNOR, WANG & XIE, 2014 (Type)
OOCLASS: AMPHIBIA LINNAEUS, 1758
OOSUBCLASS: LABYRINTHODONTIA OWEN, 1860
Incertae sedis
Oogenus: Mazonova GODFREY, 1995
M. helmichnus GODFREY, 1995 (Ootype)
OOCLASS: REPTILIA LAURENTI, 1768
OOSUBCLASS: CHELONOMORPHA KUHN, 1960
OORDER: TESTUDINES NOVA
(Eggs with testudoid basic shell type)
Family: ADOCIDAE Cope, 1870
Genus: Adocus COPE, 1870
A. sp
Family: TESTUDOOLITHIDAE Hirsch, 1996 (non FANG, LU,
JIANG & YANG, 2003)
= oofamily: CROCODILOOLITHIDAE Fang, In Fang, Zhang, Pang,
Li, Han, Xie, Yan, Pang Lu & Cheng, 2007
Note: There is no oogenus Crocodiloolithes making this name is invalid
Oogenus: Chelonoolithus KOHRING, 1998
C. braemi KOHRING, 1998 (Ootype)
Oogenus: Emydoolithus WANG,
WANG, ZHAO, ZHANG, & JIANG, 2013
E. laiyangensis WANG, WANG, ZHAO, ZHANG, & JIANG, 2013 (Type)
Oogenus: Neixiangoolithus FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG
LU & CHENG, 2007
N. yani FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG LU & CHENG,
2007 (ootype)
Oogenus: Testudoflexolithus HIRSCH,
1996
T.
bathonicae (BUCKMAN, 1859) HIRSCH, 1996 (Ootype)
= Oolithes bathonicoe BUCKMAN, 1859
Oogenus: Testudoolithus HIRSCH, 1996
= Haininchelys SCHLEICH, KÄSTLE, & GROESSENS-VAN DYCK, 1988
= Tiantaioolithus FANG, LIWU, YANGEN & LIANGFENG, 2003
T. curiosa (SCHLEICH, KÄSTLE, & GROESSENS-VAN DYCK, 1988) CHOI, KIM, PAIK, PARK, JUNG & XU, 2023
= Haininchelys curiosa SCHLEICH, KÄSTLE, & GROESSENS-VAN DYCK, 1988
T. magnirigidus ZELENITSKY, 1995 (nomen nudum)
T. hirschi KOHRING,
1999
(not
ootype)
T. jiangi (FANG, LIWU, YANGEN & LIANGFENG,
2003) JACKSON, JIN, VARRICCHIO, AZUMA & JIANG, 2008
= Tiantaioolithus jiangi FANG, LIWU, YANGEN & LIANGFENG,
2003
T. sp
Oogenus: Nova? O’BRIEN, 1995
PAN-PODOCNEMIDIDAE
"oofamily: CROCODILOOLITHIDAE"
Soft-shelled eggs of lizards and snakes
(Squamata)
Eggs with gekoid basic shell type
(Gekkota)
SUPERORDER: LACERTILIA OWEN, 1842
OORDER: SAURIA MACARTENEY, 1802 (SENSU RIEPPEL & DEBRAGA, 1996)
oofamily: Incerate sedis
Oogenus: Antarcticoolithus LEGENDRE, RUBILAR-ROGERS, MUSSER, DAVIS, OTERO, VARGAS & CLARKE, 2020
A. bradyi LEGENDRE, RUBILAR-ROGERS, MUSSER, DAVIS, OTERO, VARGAS & CLARKE, 2020 (Type)
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
= INFRAORDER: ANGUIMORPHA FURBRINGER, 1900
= PARVORDER: ANGUIMORPHA FURBINGER, 1900
Incertae sedis
SUPERCOHORT CHORISTODERA NOVA
ORDER: CHORISTODERA COPE, 1876
Family: HYPHALOSAURIDAE GAO & FOX, 2005
Genus: Hyphalosaurus GAO,
TANG & WANG, 1999
H.
baitaigouensis JI, JI, CHENG, YOU, LÜ & YUAN, 2004
OORDER: CROCODYLINES NOVA
INFRAORDER: NOTOSUCHIA GASPARINI, 1971
Incertae sedis
Genus: Yacarerani NOVAS, PAIS, POL, CARVALHO, SCANFERLA, MONES & RIGLOS,
2009
Y. boliviensis NOVAS, PAIS, POL, CARVALHO, SCANFERLA, MONES & RIGLOS,
2009
oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996
Oogenus: Bauruoolithus OLIVEIRA, SANTUCCI, ANDRADE, FULFARO, BASILIO, & BENTON,
2011
B. fragilis OLIVEIRA, SANTUCCI, ANDRADE, FULFARO, BASILIO, & BENTON, 2011
(Ootype)
Oogenus: Krokolithes HIRSCH, 1985
K. wilsoni HIRSCH, 1985 (Ootype)
K. helleri KOHRING & HIRSCH,
1996
K. dinophilus RUSSO, MATEUS, MARZOLA & BALBINO, 2017
K. sp
Oogenus: Suchoolithus RUSSO, MATEUS, MARZOLA & BALBINO, 2017
S. portucalensis RUSSO, MATEUS, MARZOLA & BALBINO, 2017 (Type)
PTEROSAURIA?
Incertae sedis
SUBORDER: RHAMPHORHYNCHOIDEA PLIENINGER, 1901
Family: WUKONGOPTERIDAE Wang, Kellner, Jiang & Meng, 2009
Genus: Kunpengopterus sp LU, UNWIN, JIN, LIU, & JI, 2009/WANG, KELLNER, CHENG JIANG, WANG, SAYAO, RODRIGUES, COSTA, LI, MENG, & ZHOU, 2015
SUBORDER: PTERODACTYLOIDEA PLIENINGER, 1901
Superfamily: CTENOCHASMATOIDEA Nopcsa, 1928 (sensu Unwin, 1995)
= LOPHOCRATIA UNWIN, 2003
Family: PTERODAUSTRIIDAE Bonaparte, 1971
Genus: Pterodaustro BONAPARTE,
1970
P. guinazui BONAPARTE, 1970 (Type)
Superfamily: PTERANODONTOIDEA Marsh, 1876 (sensu KELLNER, 1996)
Incertae sedis
Genus: Hamipterus WANG,
KELLNER, JIANG, WANG, MA, PALDOULA, CHENG, RODRIGUES, MENG, ZHANG, LI & ZHOU,
2014
H. tianshanensis WANG, KELLNER, JIANG, WANG, MA, PALDOULA, CHENG,
RODRIGUES,
MENG, ZHANG, LI & ZHOU, 2014 (Type)
OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
= GRUMULIOVUM VIALOV, 1986
= ORNATIOVUM VIALOV, 1986
Oofamily: Indet
Oogenus: Mosaicoolithus WANG, ZHAO, WANG & JIANG, 2011
M. hangtoucaoensis (FANG, WANG & JIANG, 2000) WANG, ZHAO, WANG & JIANG,
2011 (Ootype)
= Spheroolithus zhangtoucaoensis FANG, WANG & JIANG, 2000
= Spheroolithus jincunensis FANG, LU, JIANG & YANG, 2003
Oogenus: Stillatuberoolithus OSER, CHIN, SERTICH, VARRICCHIO, CHOI & RIFKIN, 2021
S. storrsi OSER, CHIN, SERTICH, VARRICCHIO, CHOI & RIFKIN, 2021 (Type)
oofamily: DINOSAURIOVIDAE Vialov, 1986
Oogenus: Dinosauriovum VIALOV, 1975
Oosubgenus: Disparovum VIALOV, 1986
D. D. compositium VIALOV, 1986 (Ootype)
Oosubgenus: Grumuliovum VIALOV, 1986
D. G. tuberculatum VIALOV, 1986 (Ootype)
D. G. asperum VIALOV, 1986
D. G. punctatum VIALOV, 1986
Oosubgenus: Laeviovum VIALOV, 1986
D. L. laevis VIALOV, 1986 (Ootype)
Oosubgenus: Ornatiovum VIALOV, 1986
D. O. multifarium VIALOV, 1986 (Ootype)
D. O. partentosum VIALOV, 1986
D. O. subtilis VIALOV, 1986
Oofamily: UMBELLAOOLITHIDAE Huang, Wang, Hu, Ren, & Wang, 2017
Oogenus: Umbellaoolithus HUANG, WANG, HU, REN, & WANG, 2017
U. xiuningensis HUANG, WANG, HU, REN, & WANG, 2017 (Ootype)
Oofamily: STALICOOLITHIDAE Wang, Wang, Zhao & Jiang, 2012
Oogenus: Coralloidoolithus WANG, WANG, ZHAO & JIAN, 2012
C. shizuiwanensis (FANG, LU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998) WANG, WANG, ZHAO & JIAN, 2012 (Ootype)
=Paraspheroolithus shizuiwanensis FANG, LU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU,
XIE & JIN, 1998
C. sp
Oogenus: Shixingoolithus ZHAO,
YE, LI, ZHAO, & ZHANG, 1991
S. erbeni ZHAO, YE, LI, ZHAO, & ZHANG,
1991 (Ootype)
Oogenus: Stalicoolithus WANG, WANG, ZHAO & JIAN, 2012
S. shifengensis WANG, WANG, ZHAO & JIAN, 2012 (Ootype)
OOCOHORT: THEROPODA NOVA
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: SPINOSAURIA OLSHEVSKY, 1991
Superfamily: MEGALOSAUROIDEA Huxley, 1889
Family: MEGALOSAURIDAE Huxley, 1889
Subfamily: MEGALOSAURINAE Huxley, 1889 (sensu Paul, 1988a)
Genus: Torvosaurus sp
EGGSHELLS OF THE ORNITHOID BASIC TYPE THEROPODA
Incertae sedis
MICRORDER: COELUROSAURIA HUENE, 1914
Family: COMPSOGNATHIDAE Cope, 1875
Genus: Compsognathus WAGNER, 1859
C. longipes WAGNER, 1859
Genus: Sinosauropteryx JI QIANG et. al., 1996
S. prima JI QIANG, et al., 1996
CLADE NAME TYRANNORAPTORA SERENO, 1999
CLADE NAME: PANTYRANNOSAURIA DELCOURT & GRILLO, 2018
CLADE NAME: EUTYRANNOSUARIA DELCOURT & GRILLO, 2018
Superfamily: TYRANNOSAUROIDEA Osborn, 1906 (sensu Bonaparte ET AL, 1990)
Family: TYRANNOSAURIDAE Osborn, 1906
Subfamily: TYRANNOSAURINAE Osborn, 1906
Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)
AVETHEROPODA PAUL, 2002
ORDER: COELURIA MARSH, 1881
SUBORDER: SEGNOSAURIA BARSBOLD & PERLE, 1980
Family: THERIZINOSAURIDAE Maleev, 1954
Oogenus: Nova? MANNING, JOYSEY, & CRUICKSHANK, 1997, 2000
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Incertae sedis
CLADE NAME: MANIRAPTOROMORPHA CAU, 2018
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: OVIRAPTOROSAURIA BARSBOLD, 1976
Superfamily: CAENAGNATHOIDEA Lamanna,Sues, Schachner, & Lyson, 2014
Family: CAENAGNATHIDAE R. M. Sternberg, 1940
Geus: Beibelong PU, ZELENITSKY, LÜ, CURRIE, CARPENTER, XU, KOPPELHUS, JIA, XIAO, CHUANG, LI, KUNDRAT & SHEN, 2017
B. sinensis PU, ZELENITSKY, LÜ, CURRIE, CARPENTER, XU, KOPPELHUS, JIA, XIAO, CHUANG, LI, KUNDRAT & SHEN, 2017 (Type)
oofamily: BELONOOLITHIDAE Jackson & Varricchio, 2016
Oogenus: Belonoolithus JACKSON & VARRICCHIO, 2016
B. garbani JACKSON & VARRICCHIO, 2016 (Ootype)
oofamily: TUBERCULOOLITHIDAE Jackson & Varricchio, 2016
Oogenus: Dimorphoolithus JACKSON & VARRICCHIO, 2016
D. bennetti JACKSON & VARRICCHIO, 2016 (Ootype)
Oogenus: Tubercuoolithus JACKSON & VARRICCHIO, 2010
T. tetonensis JACKSON & VARRICCHIO, 2010 (Ootype)
oofamily: ELONGATOOLITHIDAE Zhao, 1975
Note: Elongatoolithids are known to belong to theropod dinosaurs based on new material collected in Mongolia and China. The well-preserved remnants of an oviraptorid embryo have been found within an Elongatoolithus egg (Norell et al, 1994). Oviraptorosaur embryos have also been found in Macroelongatoolithus eggs from Henan Province, China (Zelenitsky, pers. comm.). Additionally, adult specimens of Oviraptor have been found atop of nests of Elongatoolithus eggs (Dong and Currie, 1996; Norell et al., 1995). Elongatoolithid eggshells are also known from Lower Cretaceous strata of North America (Zelenitsky pers. comm.); These rocks also contain the remains of probable Oviraptorosaurs (Makovicky and Sues, 1998).
Oogenus: Ellipsoolithus MOHABEY, 1998
E. khedaensis MOHABEY, 1998 (Ootype)
Oogenus: Elongatoolithus ZHAO, 1975
E. elongatus (YOUNG, 1954) ZHAO, 1975 (Ootype)
= Oolites elongatus YOUNG, 1954
= Spheroolithus
elongatus (YOUNG, 1954)
E. andrewsi ZHAO,
1975
E. magnus ZENG & ZHANG,
1979
E. frustrabilis MIKHAILOV,
1994
E. subtitectorius MIKHAILOV,
1994
E. sigillarius MIKHAILOV,
1994
E. excellens MIKHAILOV,
1994
E. chichengshanensis FANG, LU, JIANG & YANG, 2003
E. laijiaensis FANG, LU, JIANG & YANG, 2003
E. jianchangensis FANG, in FANG, ZHANG,
PANG, LI, HAN, XIE, YAN, PANG LU & CHENG,
2007
E. yangjiagouensis FANG, in FANG, ZHANG,
PANG, LI, HAN, XIE, YAN, PANG LU & CHENG,
2007
E. chimeiensis FANG, in FANG, ZHANG,
PANG, LI, HAN, XIE, YAN, PANG LU & CHENG,
2007
E. sp
Elongatolithus?
Genus: Citipati CLARK, NORELL & BARSBOLD, 2001
C. osmolskae CLARK, NORELL & BARSBOLD, 2001Genus: Oviraptor OSBORN, 1924
O. philoceratops OSBORN, 1924Genus: Ingenia BARSBOLD, 1981
Sp
Oogenus: Heishanoolithus ZHAO & ZHAO, 1999
H. changii ZHAO & ZHAO, 1999 (Ootype)
Oogenus: Lepidotoolithus ZHANG & XUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN,
1996)
L. guofenglouensis ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN,
1996) (Ootype)
Oogenus: Macroelongatoolithus LI, AN, ZHU, ZHANG, LIU, QU, YOU, LIANG,
LI, et al.,1995
M. carlylei (JENSEN, 1970) ZELENITSKY, CARPENTER & CURRIE, 2000
= Oolithes carlyensis JENSEN, 1970
= Oolithes carliensis MIKHAILOV, 1991 (sic)
= Spheroolithus carlyensis (JENSEN, 1970)
= Boletuoolithis carlyensis (JENSEN,
1970) ZELENITSKY, CARPENTER & CURRIE,
2000
M. xixiaensis LI, AN, ZHU, ZHANG,
LIU, QU, YOU, LIANG, LI, et al.,1995 (Ootype)
= Longiteresoolithus xixiaensis WANG & ZHOU, 1995
= Macroelongatoolithus xixia CARPENTER, 1999
M. goseongensis KIM, YANG, CHOI, SEA & KIM, 2010
M. qiaoxianensis WANG, ZHAO, WANG, JIANG & ZHANG, 2010
M. nova HODGES & ZANNO, 2023
Oogenus: Macroolithus ZHAO, 1975
M. rugustus (YOUNG, 1965) ZHAO, 1975 (Ootype)
= Oolithes rugustus YOUNG, 1965
M. yaotunensis ZHAO, 1975
M. mutabilis MIKHAILOV, 1994
M. sp
Oogenus: Nanhsiungoolithus ZHAO, 1975
N. chuetienensis ZHAO, 1975 (Ootype)
Oogenus: Paraelongatoolithus WANG, WANG, ZHAO & JIANG, 2010
P. reticulates WANG, WANG, ZHAO & JIANG, 2010 (Ootype)
Oogenus: Spongioolithus BRAY, 1999
S. hirschi BRAY, 1999 (Ootype)
Oogenus: Trachoolithus MIKHAILOV, 1994
T. faticanus MIKHAILOV, 1994 (Ootype)
Oogenus: Undulatoolithus WANG, ZHAO, WANG, LI & ZOU, 2013
U. pengi WANG, ZHAO, WANG, LI & ZOU, 2013 (Type)
Family: OVIRAPTORIDAE Barsbold, 1976
Oofamily: MONTANOOLITHIDAE Zelenitsky & Therrien, 2008
Oogenus: Montanoolithus ZELENITSKY & THERRIEN, 2008
M. strongorum ZELENITSKY & THERRIEN, 2008 (Ootype)
INFRAORDER: PARAVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Oogenus: Reticuloolithus ZELENITSKY & SLOBODA, 2005
R. hirschi ZELENITSKY & SLOBODA, 2005 (ootype)
Subfamily: VELOCIRAPATORINAE Barsbold, 1978
Eggshells associated with Deinonychus antirrhopus (MAKOVICKY & GRELLET-TINNER, 2000)
Family: TROODONTIDAE Gilmore, 1924
oofamily: OBLONGOOLITHIDAE Mikhailov, 1996
Oogenus: Oblongoolithus MIKHAILOV, 1996
O. glaber MIKHAILOV, 1996 (Ootype)
SUBORDER: METORNITHES PERLE, NORELL, CHIAPPE & CLARK, 1993
INFRAORDER: ALVAREZSAURIA BONAPARTE, 1991
Family: ELOPTERYGIDAE Lambrecht, 1933
Subfamily: PATAGONYKINAE Agnolin, Powelll, Novas, & Kundrat, 2012
Genus: Bonapartenykus AGNOLIN, POWELL, NOVAS & KUNDRAT, 2012
B. ultimus AGNOLIN, POWELL, NOVAS & KUNDRAT, 2012 (Type)
EGGS WITH DINOSAUROID-PRISMATIC BASIC SHELL TYPE
oofamily: PRISMATOOLITHIDAE Hirsch, 1994
Note: Prismatoolithid eggs from the Upper Cretaceous of Montana were initially attributed to hypsilophodontid dinosaurs, based on embryonic remains within eggs (Horner and Weishampel, 1888; Hirsch and Quinn, 1990). Subsequently, numerous prismatoolithid egg fossils from the Santonian-Campanian of Mongolia (oogenus Protoceratopsidovum) were attributed to protoceratopsian dinosaurs, based on the common occurrence of protoceratopsids in these strata (Mikhailov, 1994a; Mikhailov et al, 1994). It has been announced recently that assignment of the embryonic remains from the prismatoolithid eggs from Montana to the hypsilophodontid Orodromeus makelai was erroneous. The embryo has since been reassigned to the theropod Troodon cf. formosus (Horner and Weishampel 1996), Prismatoolithid eggs share several characteristics with other theropod eggs (e.g., elongatoolithids) and their initial assignment to hypsilophodonts and protoceratopsids is probably incorrect (Zelenitsky, pers. comm.).
Oogenus: Oolithus (Oolithes) non BUCKMAN, 1859
O. laminadermus CHAO & CHANG, 1974
Oogenus: Preprismatoolithus ZELENITSKY & HILLS, 1996
= Prismatoolithus HIRSCH, 1994
P. coloradensis (HIRSCH, 1994) ZELENITSKY & HILLS, 1996 (Ootype)
= Prismatoolithus coloradensis HIRSCH, 1994
P. sp
Oogenus: Prismatoolithus ZHAO & LI, 1993
P. gebiensis ZHAO & LI, 1993 (Ootype)
P. tenuis VIANEY-LIAUD & CROCHET, 1993
P. matellensis VIANEY-LIAUD & CROCHET, 1993
P. levis ZELENITSKY & HILLS,
1996
P. hanshuiensis ZHOU, REN, XU & GUAN, 1998
P. jenseni BRAY, 1999
P. hukouensis ZHAO, 1999
P. caboti GGARCIA, FEIST, CABOT, VALENTIN & VIANEY-LIAUD, 2000
P. hirschi JACKSON & VARRICCHIO, 2010
P. tiantaiensis WANG, ZHAO, WANG & JIANG, 2011
= Elongatoolithus tiantaiensis FANG, WANG & JIANG, 2000
P.
Oospecies: Nova VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
P. sp
Oogenus: Protoceratopsidovum MIKHAILOV, 1994
P. sincerum MIKHAILOV, 1994 (Ootype)
P. minimum MIKHAILOV, 1994
P. fluxuosum MIKHAILOV, 1994
Oogenus: Sankofa LOPEZ-MARTINEZ & VICENS, 2012
S. pyrenaica LOPEZ-MARTINEZ & VICENS, 2012 (Type)
Oogenus: Spheruprismatoolithus BRAY, 1999
S. condensus BRAY, 1999 (Ootype)
MORPHOTYPE: ORNITHOID-PRISMATIC INCERTAE SEDIS
Oogenus: Dispersituberoolithus ZELENITSKY, HILLS & CURRIE, 1996
D. exilis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Oogenus: Pseudogeckoolithus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
P. nodosus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997 (Ootype)
Note: Morphotype: Geckonoid?
P. sp
OOCOHORT: ORNITHISCHIFORMES NOVA
OOSUPERORDER: SAUROPODOMORPHA NOVA
Incertae sedis
Genus: Qianlong HAN, YU, ZHANG, ZENG, WANG, CAI, WU, WEN, CAI, LI, WU, ZHAO & XU, 2023
Q. shouhu HAN, YU, ZHANG, ZENG, WANG, CAI, WU, WEN, CAI, LI, WU, ZHAO & XU, 2023 (Type)
SUBORDER: PLATEOSAURIA TORNIER, 1913
ORDER: PROSAUROPODA HUENE, 1920 (emend)
SUBORDER: PLATEOSAURIA TORNIER, 1913
Family: PLATEOSAURIDAE Marsh, 1895
Genus: Mussaurus BONAPARTE & VINCE,
1979
M. patagonicus BONAPARTE & VINCE, 1979
(Type)
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
Family: MASSOSPONDYLIDAE Huene, 1914
Genus: Massospondylus OWEN,
1854
M. sp
TUBOSPHERULITIC STRUCTURAL MORPHOTYPE
oofamily: "MEGALOOLITHIDAE" Zhao, 1979
[Sauropoda]
Note: The first dinosaur eggs remains were discovered in Cretaceous strata of Southern France over a century ago (Pouech..). These eggs, known as megaloolithids, have been assumed to belong to sauropod dinosaurs because they are usually found in strata that are rich in sauropod remains (Erben et al., 1979; Vianey-Liaud et al. 1987; Jain 1989; Sahni et al, 1994). Recent confirmation of this correlation was made with the discovery of South American megaloolithid eggs containing sauropod embryos (Chiappe et al, 1998). Occasional references to typical megaloolithid eggshells as “hadrosaurian’ (Grigorescu et al, 1994, p. 77) are obviously misleading. Additionally, Upper Cretaceous strata in North America are rich hadrosaur remains and contain no trace of megaloolithid eggshell (Zelenitsky, pers. comm.). All positively identified hadrosaurian eggs have a different eggshell structure (see Spheroolithidae).
Oogenus: Cairanoolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
= Dughioolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
C. dughii VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994 (Ootype)
C. roussetensis (VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994) BARCIA & VIANEY-LIAUD,
2001
= Dughioolithus roussetensis VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD,
1994
Oogenus: Fusioolithus
F.
F. sp
Oogenus: Megaloolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. mammilare VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994 (Ootype)
M. siruguei VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. aurelinesis VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. petralta VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. cylindricus KHOLSA & SAHNI, 1995
= Megaloolithus rahioliensis MOHABEY, 1998
M. jabalpurensis KHOLSA & SAHNI, 1995
= Megaloolithus patagonicus CALVO, ENGLLAND, HEREDIA & SALGADO, 1997
= Megaloolithus matleyi MOHABEY, 1997
M. mohabeyi KHOLSA & SAHNI, 1995
= Megaloolithus phensaniensis MOHABEY, 1998
M. dholiyaensis KHOLSA & SAHNI, 1995
M. padiyalensis KHOLSA & SAHNI, 1995
M. walpurensis KHOLSA & SAHNI, 1995
= Megaloolithus khempurensis MOHABEY, 1998
Note: VIANEY-LIAUD, KHOSLA & GARCIA, 2003 place Megaloolithus walpurensis within the M.
khempurensis because of the fragmentary nature of M. walpurensis,
but M. walpurensis has priority.
M.
trempii MORATALLA, 1998
M. dhoridungriensis MOHABEY,
1998
M. megadermus MOHABEY,
1998
M. microtuberculata GARCIA & VIANEY-LIAUD,
2001
M. maghrebiensis GARCIA, TABUCE, CAPPETTA, MARANDAT, BENTALEB, BENABDALLAH, & VIANEY-LIAUD,
2003
M. nova PANDES
I BLAS, 2005
M. sp
Incertae sedis
Oogenus: Tacuaremborum MONES, 1980
T. oblongum MONES, 1980 (Ootype)
Oofamily: POLYCLONOOLITHIDAE Xie, Zhang, Jin, Li, & Zhou, 2015
Genus: Polyconoolithus XIE, ZHANG, JIN, LI, & ZHOU, 2015
P. yangjiagouensis
XIE, ZHANG, JIN, LI, & ZHOU, 2015 (Type)
oofamily: FAVEOLOOLITHIDAE Zhao & Ding, 1976 [Sauropoda]
Note: MIKHAILOV, 1997…Only bones of sauropod dinosaurs are known from the Faveoloolithus nest-bearing layers in Ologoy-Ulan-Tsav locality in Mongolia Gobi (Sochava, 1969; S. M. Kurzanov, pers. comm.). The arrangement of the eggs and porosity of the eggshell appear to be similar to those subterranean nesters, such as chelonian.
Oogenus: Faveoloolithus ZHAO & DING, 1976
F. ningxiaensis ZHAO & DING, 1976 (Ootype)
F. sp
Oogenus: Hemifaveoloolithus WANG, ZHAO, WANG & JIANG, 2011
H muyushanensis WANG, ZHAO, WANG & JIANG, 2011 (Ootype)
Oogenus: Parafaveoloolithus ZHANG, 2010
P. microporus ZHANG, 2010 (Ootype)
P. macroporus ZHANG, 2010
P. tiansicunensis ZHANG, 2010
P. guoqingsiensis (FANG, WANG & JIANG, 2000) WANG, ZHAO, WANG & JIANG,
2011
= Dendroolithus guoqingsiensis FANG, WANG & JIANG,
2000
P. pingxiagensis ZOU, WANG & WANG, 2013
Oogenus: Sphaerovum MONES, 1980
S. erbeni MONES, 1980 (Ootype)
S. sp
Oofamily: FUSIOOLITHIDAE Fernandez & Kholsa, 2014
Oogenus: Fusioolithus FERNANDEZ & KHOSLA, 2014
= Patagoolithus SIMON, 2006
F. baghensis (KHOLSA & SAHNI, 1995) FERNANDEZ & KHOSLA, 2014 (Ootype)
= Megaloolithus baghensis KHOLSA & SAHNI, 1995
= Megaloolithus pseudomamillareVIANEY-LIAUD, HERISCH, SAHNI & SIGE, 1997
= Megaloolithus balasinorensis MOHABEY, 1998
= Patagoolithus salitralensis SIMON, 2006
F. berthei FERNANDEZ & KHOSLA, 2014
Oofamily: SIMILIFAVEOLOOLITHIDAE Wang, Zhao,Wang & Jiang, 2011
Oogenus: Simlifaveoloolithus WANG, ZHAO, WANG & JIANG, 2011
S. shuangtangensis (FANG, LU, JIANG & YANG, 2003) WANG, ZHAO, WANG & JIANG,
2011 (Ootype)
= Dendroolithus shuangtangensis FANG,
LU, JIANG & YANG, 2003
oofamily: YOUNGOOLITHIDAE Zhang, 2010
Oogenus: Youngoolithus ZHAO 1979
Y. xiaguanensis ZHAO, 1979(Ootype)
Y. xipingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU,
XIE & JIN, 1998
Y. sp
Family: Unknown.
Family: Titanosauridae?
oofamily: DENDROOLITHIDAE Zhao and Li, 1988
[Sauropoda? Therizinosauria? Mikhailov, 1997
Note: As per Mikhailov, 1997…Based on the taphonomic settings and reconstructed type of nesting, dendroolithid remains have been attributed tentatively to some ornithopod or sauropod dinosaurs (Mikhailov, 1991). Recently, some privately owned dendroolithid eggs from Henan Province, China displayed at the University Museum of Zoology, Cambridge (the author was kindly allowed to examine shell samples) have been shown to contain well preserved embryos of therizinosaurid dinosaurs at late stages of development (Cohen et al, 1995)…
Oogenus: Dendroolithus ZHAO & LI, 1988
D. wangdianensis ZHAO & LI, 1988 (Ootype)
D. verrucarius MIKHAILOV, 1994
D. microporosus MIKHAILOV, 1994
D. xichuanensis ZHAO & ZHAO, 1998
D. furcatus FANG, LU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN,
1998
D. dendriticus FANG, LU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN,
1998
D. sanlimiaoensis FANG, LU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN,
1998
D. zhaoyingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU,
XIE & JIN, 1998
D. hongzhaiziensis ZHOU, REN, XU & GUAN, 1998
D. tumiaolingensis ZHOU, REN, XU & GUAN, 1998
D. fengguangcunensis FANG, ZHANG, ZHANG, LU, HAN & LI, 2005
D. dashiqiaoensis JIA, FANG, GAO, GAO, CHANG, CHANG, XIAO, YANG, ZHENG, ZHU, WANG & WANG, 2020
D. sp
Oogenus: Placoolithus ZHAO & ZHAO,
1998
= Paradendroolithus ZHOU, REN, XU & GUAN, 1998
P. taohensis ZHAO & ZHAO, 1998 (Ootype)
P. tumiaolingensis (ZHOU, REN, XU & GUAN, 1998) ZHANG, YANG, LI & HU, 2018
= Dendroolithus tumiaolingensis ZHOU, REN, XU & GUAN, 1998
= Dendroolithus hongzhaiziensis ZHOU, REN, XU & GUAN, 1998
= Paradendroolithus ginglongshanensis ZHOU, REN, XU & GUAN, 1998
Oogenus: Prionoolithus HE, ZHANG, XING, JIANG, AN & YANG, 2019
P. quyuangangensis HE, ZHANG, XING, JIANG, AN & YANG, 2019 (Ootype)
oofamily: DICTYOOLITHIDAE Zhao, 1994
Oogenus: Dictyoolithus ZHAO, 1994
D. hongpoensis ZHAO, 1994 (Ootype)
Oogenus: Paradictyoolithus WANG, ZHAO, WANG, ZHANG, & JIANG,
2013
P. zhuangqianensis WANG, ZHAO, WANG, ZHANG, & JIANG, 2013 (Ootype)
P. xiaxishanensis WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
Oogenus: Protodictyoolithus WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
P. neixiangensis (ZHAO, 1994) WANG, ZHAO, WANG, ZHANG, & JIANG, 2013 (Ootype)
= Dictyoolithus neixiangensis ZHAO, 1994
P. jiangi (LIU & ZHAO, 2004) WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
= Dictyoolithus jiangi LIU& ZHAO, 2004
Oogenus: Stromatoolithus ZHAO,
YE, LI, ZHAO & YAN, 1991
S. pinglingensis ZHAO, YE, LI, ZHAO & YAN,
1991 (Ootype)
S. gongzhuligensis (WANG, ZAN, JIN & CHEN, 2005) WANG, ZHAO, WANG, ZHANG, & JIANG,
2013
= Dictyoolithus gongzhuligensis WANG, ZAN, JIN & CHEN, 2005
oofamily: PHACELOOLITHIDAE Zeng& Zhang, 1979
Oogenus: Phaceloolithus ZENG & ZHANG, 1979
P. huanensis ZENG & ZHANG, 1979 (Ootype)
Gen. sp indet.
oofamily: Incertae sedis ?spherulite or spheruprismatic structural type eggshell.
Oogenus: Parvoolithus MIKHAILOV, 1996
P. tortuosus MIKHAILOV, 1996 (Ootype)
Note: Possible beetle castings ?
Gen. sp indet.
oofamily: Incertae sedis
Oogenus: Placoolithus non ZHAO, 1979
P. cf. taohensis ZHAO, 1979 (Ootype)
OOMAGNORDER: ORNITHISCHIA SEELEY, 1888
EGGS WITH DINOSAUROID-SPHERULITIC BASIC SHELL TYPE
oofamily: DONGYANOOLITHIDAE Zhang, Xie, Jin, Du, & Huang, 2019
Oogenus: Dongyangosaurus JIN, 2013
D. sinensis JIN, 2013 (Ootype)
D. namaensis
Oogenus: Multifissolithus ZHANG, XIE, JIN, DU & HUANG, 2019
M. chianensis ZHANG, XIE, JIN, DU & HUANG, 2019 (Ootype)
oofamily: SPHEROOLITHIDAE Zhao, 1979
[Ornithopoda: some hadrosaurs]
Note: As per Mikhailov, 1997…Spheroolithus eggs from Laiyang have been attributed to hadrosaurs (Young, 1959, pp. 511, 517); the eggs of S. irenensis were found in close association with Bactrosaurus remains (Zhao and Jiang, 1974, p. 81). Spheroolithid eggs from the Upper Cretaceous Two Medicine Formation of Montana were found in association with hadrosaurian juveniles (Horner and Makela, 1979). Lambeosaurine embryos (Hypacrosaurus stebingeri; Horner and Currie, 1994) were also found in association with large spherical eggs; however, the eggshell morphotype could not be identified as spheroolithid because it is diagenetically modified (Zelenitsky, 1995). This was misinterpreted by Mikhailov. These discoveries have allowed all spheroolithid remains to be attributed to hadrosaurs (Hirsch and Quinn, 1990, Mikhailov et al, 1994). Their abundance, as well as their geographical and stratigraphical distribution, are in accordance with this conclusion…
Oogenus: GuegoolithusMORENO-AZANZA, CANUDO & GASCA, 2014
G. turolensis (AMO SANJUAN, CANUDO & CUENCA BESCOS, 2000) MORENO-AZANZA, CANUDO & GASCA, 2014 (Ootype)
=Macroolithus turolensis AMO SANJUAN, CANUDO & CUENCA BESCOS, 2000
Oogenus: Paraspheroolithus ZHAO, 1979
Note: (= Spheroolithus ZHAO, 1979 as per MIKHAILOV, 1997)
P. irenensis (CHAO & CHIANG, 1974) ZHAO, 1979 (Ootype)
= Spheroolithus irenensis (CHAO & CHIANG, 1974) MIKHAILOV, 1994
= Oolithes spheroides YOUNG, 1954
= Oolithes irenensis CHAO & CHIANG, 1974
= Spheroolithus sp MIKHAILOV, 1991
P. lamelliformae ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN,
1996)
P. yangchengensis FANG, LU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU,
XIE & JIN, 1998
P. sanwangbacunensis FANG, ZHANG, ZHANG, LU, HAN & LI, 2005
P. sp
Oogenus: Spheroolithus ZHAO, 1979
S. spheroides (YOUNG, 1954) ZHAO, 1979
(Ootype)
= Oolithes spheroides YOUNG, 1954
S. chiangchiungtingensis (CHAO & CHIANG,
1974) ZHAO, 1979
= Oolithes chiangchiungtigensis CHAO & CHIANG,
1974
S. megadermus (YOUNG, 1959) ZHAO, 1979
= Oolithes megadermus YOUNG, 1959
S. tenuicorticus MIKHAILOV, 1994
S. maiasauroides MIKHAILOV, 1994
S. albertensis ZELENITSKY & HILLS,
1997
S. choteauensis JACKSON & VARRICCHIO, 2010
S. europaeus SELLES, VIA, & GALOBART, 2014
Oosp (unnamed)
= Genus: Maiasaura HORNER & MAKELA,
1979
M. peeblesorum HORNER & MAKELA, 1979
S. sp
Oogenus: Shangyangoolithus ZHANG & XUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN,
1996)
= Shanyangoolithus ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN,
1996) (sic)
S. shanyangensis ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN,
1996) (Ootype)
Oogenus: Nova XU, ZHAO, LU, DONG & HUANG, 1999
Incertae sedis
Oogenus: Triprismatoolithus JACKSON & VARRICCHIO, 2010
T. stephensi JACKSON & VARRICCHIO, 2010 (Ootype)
oofamily: OVALOOLITHIDAE Mikhailov, 1991
(?Ornithopoda)
Note: As per Mikhailov, 1997…The histrostructure of the shell and other, superficial features of the egg, are similar to those of the Spheroolithidae. At the Mogoyn-Ulagiyn-Kheates locality (= Tel-Ulan-Ula; Mongolia, Dornogov Aimak), pinkish strata containing numerous ovaloolithid eggs in clutches yield remnants of only lambeosaurine dinosaurs (S. M. Kurzanov, pers. comm.). This prompts the tentative assignment of ovaloolithid remains to ornithopod dinosaurs. Preliminary assignment of embryonic limb bones associated with an ovaloolithid egg (PIN 2970/9) to protoceratopsian dinosaurs (L. I. Hozatcky in Sochen, 1972) has proved to be incorrect (S. M. Kurzanov, see Mikhailov et al., 1994)…
Oogenus: Ovaloolithus ZHAO, 1978
O. chinkangkouensis (CHAO & CHIANG,
1974) ZHAO, 1978 (Ootype)
= Oolithes chinkangkouensis CHAO & CHIANG, 1974
O. laminadermus (CHAO & CHIANG, 1974) ZHAO, 1979
= Oolithes sp YOUNG, 1954
= Oolithes laminadermus (YOUNG, 1954) CHAO & CHIANG, 1974
O. tristriatus ZHAO, 1979
O. mixtistriatus ZHAO, 1979
O. monostriatus ZHAO, 1979
O. dinornithoides MIKHAILOV, 1994
O. sangpingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU,
XIE & JIN, 1998
O. utahensis BRAY, 1999
O. tenuisus BRAY, 1999
O. sp
Ornithischia incertae sedis?
HYPSILOPHODONTIDAE
RHABDODONTIDAE
DRYOSAURIDAE
CAMPTOSAURIDAE
HADROSAURIDAE
Saurolophus angustirostris (Spheroolithus)
LAMBEOSAURINAE
PSITTACOSAURIDAE
PROTOCERATOPIDAE?
CERATOPIAN
KENTROSAURINAE
OOCLASS: AVES NOVA
Incertae sedis
Oogenus: Plagioolithus IMAI & AZUMA, 2014
P. fukuiensis IMAI & AZUMA, 2014 (Ootype)
COHORT: ENANTIORNITHES NOVA
= SUBCLASS: ENANTIORNITHES WALKER, 1981
= INFRACLASS: ENANTIORNITHES WALKER,
1981
Note: = AERIALAE JI & JI, 2001 (partim)
Incertae sedis
Genus: Avimaia BAILLEUL, O’CONNOR, ZHANG, LI, WANG, LAMANNA, ZHU & ZHOU, 2019
A. schweitzerae BAILLEUL, O’CONNOR, ZHANG, LI, WANG, LAMANNA, ZHU & ZHOU, 2019 (Type)
oofamily: LAEVISOOLITHIDAE Mikhailov, 1991 [AVES, Enantiornithes]
= oofamily: SUBTILIOLITHIDAE Mikhailov, 1991 [AVES]
Note: As per Mikhailov, 1997… These small and thin-shelled eggs with ornithoid eggshell structure were proposed to belong either to birds or small theropod dinosaurs (Mikhailov, 1991). Recently, one of the specimens of laevisoolithid eggshells was found in situ with an associated partial skeleton of an enantiornithid bird Nanantius (Kurockhin, 1996). This allows taxonomic assignment of laevisoolithid eggs to Enantiornithes (Mikhailov, 1996a)…
Oogenus: Laevisoolithus MIKHAILOV, 1991
L. sochavai MIKHAILOV, 1991 (Ootype)
Genus: Gobipteryx ELZANOWSKI, 1976
G. minuta ELZANOWSKI, 1976
= Nanantius valifanovi KUROCHKIN, 1996
G. sp
Oogenus: Subtiliolithus MIKHAILOV,
1991
S. microtuberculatus MIKHAILOV, 1991
(Ootype)
S.
kachchhensis KHOLSA & SAHNI,
1995
Oogenus: Tipoolithus GARCIA, TABUCE, CAPPETTA, MARANDAT, BENTALEB, BENABDALLAH, & VIANEY-LIAUD,
2003
T. achloujensis GARCIA, TABUCE, CAPPETTA, MARANDAT, BENTALEB, BENABDALLAH, & VIANEY-LIAUD,
2003 (Ootype)
Oofamily: PACHYCORIOOLITHIDAE Lawver, Jin, Jackson & Wang, 2016
Oogenus: Pachycorioolithus LAWVER, JIN, JACKSON & WANG, 2016
P. jinyuensis LAWVER, JIN, JACKSON & WANG, 2016 (Ootype)
oofamily: GOBIOOLITHIDAE Mikhailov, 1996 [Aves]
Note: As per Mikhailov, 1997…Well-developed embryos of palaeognathous birds have been found at Khermiyn-Tsav (Elzanowski, 1981; Kurochkin, 1988). Although Elzanowski never stated that these embryos should be assigned to the small bird Gobipteryx minuta, described earlier from deposits of the same age (Elzanowski, 1977), this opinion has occasionally appeared in the literature Recent re-examination of embryonic material (Kurockhin, 1995) has shown that the embryo, which shares some features with Gobipteyx, is not certainly associated with the Gobioolithus minor eggs. Embryos that are certainly from Gobioolithus minor eggs may belong to a different group of birds, apparently Palaeognathae…
Genus: Gobipipus CHATTERJEE, 1994 (noma nudum)
G. elzanowskii CHATTERJEE, 1994 (noma nudum)
Oogenus: Gobioolithus MIKHAILOV, 1996
G. minor MIKHAILOV, 1996 (Ootype)
G.
maior MIKHAILOV, 1996
Oogenus: Styloolithus VARRICCHIO & BARTHA, 2015
S. sabathi VARRICCHIO & BARTHA, 2015 (Ootype)
oofamily: ORNITHOOLITHIDAE Fang, in Fang, Zhang, Pang, Li, Han, Xie, Yan, Pang Lu & Cheng, 2007
Oogenus: Protornithoolithus FANG,
in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG, LÜ & CHENG, 2007
P. tumendongensis FANG, in FANG,
ZHANG, PANG, LI, HAN, XIE, YAN, PANG, LÜ & CHENG,
2007 (Ootype)
Morphotype: Ornithoid-ratite
Incertae sedis
Oogenus: Ageroolithus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
A. fontllongensis VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997 (Ootype)
A. sp
Oogenus: Continuoolithus ZELENITSKY, HILLS & CURRIE, 1996
C. canadensis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably small theropod.
C. sp
Oogenus: Porituberoolithus ZELENITSKY, HILLS & CURRIE, 1996
P. warnerensis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably small theropod.
P. sp
Oogenus: Tristraguloolithus ZELENITSKY, HILLS & CURRIE, 1996
T. cracioides ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably belongs to a bird.
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_____________________________________________________________________________________
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A
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C
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