VETEROVATA
(Parataxonomy of fossil eggs and shells, nests?)

NESTS

Genus sp indet?

oofamily: Indet.

Oogenus: Oolithus (Oolithes) non BUCKMAN, 1859
O. sphaericus van STRAELEN, 1928
O. megadermus YOUNG, 1959
O. nanhsiungensis YOUNG, 1965

Gen. sp indet.

OOCLASS: AMPHIBIA LINNAEUS, 1758

OOSUBCLASS: LABYRINTHODONTIA OWEN, 1860

Incertae sedis

Oogenus: Mazonova GODFREY, 1995
M. helmichnus GODFREY, 1995 (Ootype)

Gen. sp indet.

OORDER: TESTUDINES NOVA
(Eggs with testudoid basic shell type)

Family: ADOCIDAE Cope, 1870

Genus: Adocus COPE, 1870
A. sp

Family: TESTUDOOLITHIDAE Hirsch, 1996 (non FANG, LU, JIANG & YANG, 2003)

Oogenus: Emydoolithus WANG, WANG, ZHAO, ZHANG, & JIANG, 2013
E. laiyangensis WANG, WANG, ZHAO, ZHANG, & JIANG, 2013 (Type)

Oogenus: Testudoflexolithus HIRSCH, 1996
T. bathonicae (BUCKMAN, 1859) HIRSCH, 1996 (Ootype)
= Oolithes bathonicoe BUCKMAN, 1859

Oogenus: Testudoolithus HIRSCH, 1996
= Tiantaioolithus FANG, LIWU, YANGEN & LIANGFENG, 2003
T. hirschi KOHRING, 1999 (not ootype)
T. jiangi (FANG, LIWU, YANGEN & LIANGFENG, 2003) JACKSON, JIN, VARRICCHIO, AZUMA & JIANG, 2008
= Tiantaioolithus jiangi FANG, LIWU, YANGEN & LIANGFENG, 2003
T. sp

Oogenus: Nova? O’BRIEN, 1995

Gen. sp indet.

Soft-shelled eggs of lizards and snakes
(Squamata)

Gen. sp indet.

Eggs with gekoid basic shell type
(Gekkota)

Gen. sp indet.

SUPERCOHORT CHORISTODERA NOVA

ORDER: CHORISTODERA COPE, 1876

Family: HYPHALOSAURIDAE GAO & FOX, 2005

Genus: Hyphalosaurus GAO, TANG & WANG, 1999
H. baitaigouensis JI, JI, CHENG, YOU, LÜ & YUAN, 2004

OORDER: CROCODYLINES NOVA

INFRAORDER: NOTOSUCHIA GASPARINI, 1971

Incertae sedis

Genus: Yacarerani NOVAS, PAIS, POL, CARVALHO, SCANFERLA, MONES & RIGLOS, 2009
Y. boliviensis NOVAS, PAIS, POL, CARVALHO, SCANFERLA, MONES & RIGLOS, 2009

oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996

Oogenus: Bauruoolithus OLIVEIRA, SANTUCCI, ANDRADE, FULFARO, BASILIO, & BENTON, 2011
B. fragilis OLIVEIRA, SANTUCCI, ANDRADE, FULFARO, BASILIO, & BENTON, 2011 (Ootype)

Oogenus: Krokolithes HIRSCH, 1985
K. wilsoni HIRSCH, 1985 (Ootype)
K. helleri KOHRING & HIRSCH, 1996
K. sp

Gen. sp indet.

oofamily: CROCODILOOLITHIDAE Fang, In Fang, Zhang, Pang, Li, Han, Xie, Yan, Pang Lu & Cheng, 2007
Note: There is no oogenus Crocodiloolithes making this name is invalid

Oogenus: Neixiangoolithus FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG LU & CHENG, 2007
N. yani FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG LU & CHENG, 2007 (ootype)

Gen. sp indet.

PTEROSAURIA?

Incertae sedis

Gen. sp indet.

SUBORDER: RHAMPHORHYNCHOIDEA PLIENINGER, 1901

Family: WUKONGOPTERIDAE Wang, Kellner, Jiang & Meng, 2009

Genus: Darwinopterus LU, UNWIN, JIN, LIU, & JI, 2009
D. modularis LU, UNWIN, JIN, LIU, & JI, 2009

SUBORDER: PTERODACTYLOIDEA PLIENINGER, 1901

Superfamily: CTENOCHASMATOIDEA Nopcsa, 1928 (sensu Unwin, 1995)
= LOPHOCRATIA UNWIN, 2003

Family: PTERODAUSTRIIDAE Bonaparte, 1971

Genus: Pterodaustro BONAPARTE, 1970
P. guinazui BONAPARTE, 1970 (Type)

Superfamily: PTERANODONTOIDEA Marsh, 1876 (senus KELLNER, 1996)

Incertae sedis

Genus: Hamipterus WANG, KELLNER, JIANG, WANG, MA, PALDOULA, CHENG, RODRIGUES, MENG, ZHANG, LI & ZHOU, 2014
H. tianshanensis WANG, KELLNER, JIANG, WANG, MA, PALDOULA, CHENG, RODRIGUES, MENG, ZHANG, LI & ZHOU, 2014 (Type)

OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
= GRUMULIOVUM VIALOV, 1986
= ORNATIOVUM VIALOV, 1986

Oofamily: Indet

Oogenus: Mosaicoolithus WANG, ZHAO, WANG & JIANG, 2011
M. hangtoucaoensis (FANG, WANG & JIANG, 2000) WANG, ZHAO, WANG & JIANG, 2011 (Ootype)
= Spheroolithus zhangtoucaoensis FANG, WANG & JIANG, 2000
= Spheroolithus jincunensis FANG, LU, JIANG & YANG, 2003

Gen. Sp indet.

oofamily: DINOSAURIOVIDAE Vialov, 1986


Oogenus: Dinosauriovum VIALOV, 1975
Oosubgenus: Disparovum VIALOV, 1986
D. D. compositium VIALOV, 1986 (Ootype)

Oosubgenus: Grumuliovum VIALOV, 1986
D. G. tuberculatum VIALOV, 1986 (Ootype)
D. G. asperum VIALOV, 1986
D. G. punctatum VIALOV, 1986

Oosubgenus: Laeviovum VIALOV, 1986
D. L. laevis VIALOV, 1986 (Ootype)

Oosubgenus: Ornatiovum VIALOV, 1986
D. O. multifarium VIALOV, 1986 (Ootype)
D. O. partentosum VIALOV, 1986
D. O. subtilis VIALOV, 1986

OOCOHORT: THEROPODA NOVA


EGGSHELLS OF THE ORNITHOID BASIC TYPE THEROPODA

Incertae sedis

Gen. sp indet.

Family: COMPSOGNATHIDAE Cope, 1875

Genus: Compsognathus WAGNER, 1859
C. longipes WAGNER, 1859

Genus: Sinosauropteryx JI QIANG et. al., 1996
S. prima JI QIANG, et al., 1996

Family: THERIZINOSAURIDAE Maleev, 1954

Oogenus: Nova? MANNING, JOYSEY, & CRUICKSHANK, 1997, 2000

Family: ELOPTERYGIDAE Lambrecht, 1933

Subfamily: PATAGONYKINAE Agnolin, Powelll, Novas, & Kundrat, 2012

Genus: Bonapartenykus AGNOLIN, POWELL, NOVAS & KUNDRAT, 2012
B. ultimus AGNOLIN, POWELL, NOVAS & KUNDRAT, 2012 (Type)

oofamily: ELONGATOOLITHIDAE Zhao, 1975

Note: Elongatoolithids are known to belong to theropod dinosaurs based on new material collected in Mongolia and China. The well-preserved remnants of an oviraptorid embryo have been found within an Elongatoolithus egg (Norell et al, 1994). Oviraptorosaur embryos have also been found in Macroelongatoolithus eggs from Henan Province, China (Zelenitsky, pers. comm.). Additionally, adult specimens of Oviraptor have been found atop of nests of Elongatoolithus eggs (Dong and Currie, 1996; Norell et al., 1995). Elongatoolithid eggshells are also known from Lower Cretaceous strata of North America (Zelenitsky pers. comm.); These rocks also contain the remains of probable Oviraptorosaurs (Makovicky and Sues, 1998).

Oogenus: Ellipsoolithus MOHABEY, 1998
E. khedaensis MOHABEY, 1998 (Ootype)

Oogenus: Elongatoolithus ZHAO, 1975
E. elongatus (YOUNG, 1954) ZHAO, 1975 (Ootype)
= Oolites elongatus YOUNG, 1954
= Spheroolithus elongatus (YOUNG, 1954)
E. andrewsi ZHAO, 1975
E. magnus ZENG & ZHANG, 1979
E. frustrabilis MIKHAILOV, 1994
E. subtitectorius MIKHAILOV, 1994
E. sigillarius MIKHAILOV, 1994
E. excellens MIKHAILOV, 1994
E. chichengshanensis FANG, LU, JIANG & YANG, 2003
E. laijiaensis FANG, LU, JIANG & YANG, 2003
E. jianchangensis FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG LU & CHENG, 2007
E. yangjiagouensis FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG LU & CHENG, 2007
E. chimeiensis FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG LU & CHENG, 2007
E. sp

Elongatolithus?

Genus: Oviraptor OSBORN, 1924
O. philoceratops OSBORN, 1924

Genus: Ingenia BARSBOLD, 1981
Sp

Oogenus: Heishanoolithus ZHAO & ZHAO, 1999
H. changii ZHAO & ZHAO, 1999 (Ootype)

Oogenus: Lepidotoolithus ZHANG & XUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996)
L. guofenglouensis ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996) (Ootype)

Oogenus: Macroelongatoolithus LI, AN, ZHU, ZHANG, LIU, QU, YOU, LIANG, LI, et al.,1995
M. carlylei (JENSEN, 1970) ZELENITSKY, CARPENTER & CURRIE, 2000
= Oolithes carlyensis JENSEN, 1970
= Oolithes carliensis MIKHAILOV, 1991 (sic)
= Spheroolithus carlyensis (JENSEN, 1970)
= Boletuoolithis carlyensis (JENSEN, 1970) ZELENITSKY, CARPENTER & CURRIE, 2000
M. xixiaensis LI, AN, ZHU, ZHANG, LIU, QU, YOU, LIANG, LI, et al.,1995 (Ootype)
= Longiteresoolithus xixiaensis WANG & ZHOU, 1995
= Macroelongatoolithus xixia CARPENTER, 1999

Oogenus: Macroolithus ZHAO, 1975
M. rugustus (YOUNG, 1965) ZHAO, 1975 (Ootype)
= Oolithes rugustus YOUNG, 1965
M. yaotunensis ZHAO, 1975
M. mutabilis MIKHAILOV, 1994
M. turolensis AMO SANJUAN, CANUDO & CUENCA BESCOS, 2000
M. sp

Oogenus: Nanhsiungoolithus ZHAO, 1975
N. chuetienensis ZHAO, 1975 (Ootype)

Oogenus: Paraelongatoolithus WANG, WANG, ZHAO & JIANG, 2010
P. reticulates WANG, WANG, ZHAO & JIANG, 2010 (Ootype)

Oogenus: Spongioolithus BRAY, 1999
S. hirschi BRAY, 1999 (Ootype)

Oogenus: Trachoolithus MIKHAILOV, 1994
T. faticanus MIKHAILOV, 1994 (Ootype)

Oogenus: Undulatoolithus WANG, ZHAO, WANG, LI & ZOU, 2013
U. pengi WANG, ZHAO, WANG, LI & ZOU, 2013 (Type)

Gen. Sp indet.

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex

Oofamily: MONTANOOLITHIDAE Zelenitsky & Therrien, 2008

Oogenus: Montanoolithus ZELENITSKY & THERRIEN, 2008
M. strongorum ZELENITSKY & THERRIEN, 2008 (Ootype)

Family: DROMAEOSAURIDAE Russell, 1969

Oogenus: Reticuloolithus ZELENITSKY & SLOBODA, 2005
R. hirschi ZELENITSKY & SLOBODA, 2005 (ootype)

Subfamily: VELOCIRAPATORINAE Barsbold, 1978

Eggshells associated with Deinonychus antirrhopus (MAKOVICKY & GRELLET-TINNER, 2000)

Family: TROODONTIDAE Gilmore, 1924

Troodon formosus

oofamily: OBLONGOOLITHIDAE Mikhailov, 1996

Oogenus: Oblongoolithus MIKHAILOV, 1996
O. glaber MIKHAILOV, 1996 (Ootype)

EGGS WITH DINOSAUROID-PRISMATIC BASIC SHELL TYPE

Gen. sp indet.

oofamily: PRISMATOOLITHIDAE Hirsch, 1994

Note: Prismatoolithid eggs from the Upper Cretaceous of Montana were initially attributed to hypsilophodontid dinosaurs, based on embryonic remains within eggs (Horner and Weishampel, 1888; Hirsch and Quinn, 1990). Subsequently, numerous prismatoolithid egg fossils from the Santonian-Campanian of Mongolia (oogenus Protoceratopsidovum) were attributed to protoceratopsian dinosaurs, based on the common occurrence of protoceratopsids in these strata (Mikhailov, 1994a; Mikhailov et al, 1994). It has been announced recently that assignment of the embryonic remains from the prismatoolithid eggs from Montana to the hypsilophodontid Orodromeus makelai was erroneous. The embryo has since been reassigned to the theropod Troodon cf. formosus (Horner and Weishampel 1996), Prismatoolithid eggs share several characteristics with other theropod eggs (e.g., elongatoolithids) and their initial assignment to hypsilophodonts and protoceratopsids is probably incorrect (Zelenitsky, pers. comm.).

Oogenus: Oolithus (Oolithes) non BUCKMAN, 1859
O. laminadermus CHAO & CHANG, 1974

Oogenus: Preprismatoolithus ZELENITSKY & HILLS, 1996
= Prismatoolithus HIRSCH, 1994
P. coloradensis (HIRSCH, 1994) ZELENITSKY & HILLS, 1996 (Ootype)
= Prismatoolithus coloradensis HIRSCH, 1994
P. sp

Oogenus: Prismatoolithus ZHAO & LI, 1993
P. gebiensis ZHAO & LI, 1993 (Ootype)
P. tenuis VIANEY-LIAUD & CROCHET, 1993
P. matellensis VIANEY-LIAUD & CROCHET, 1993
P. levis ZELENITSKY & HILLS, 1996
P. hanshuiensis ZHOU, REN, XU & GUAN, 1998
P. jenseni BRAY, 1999
P. hukouensis ZHAO, 1999
P. caboti GGARCIA, FEIST, CABOT, VALENTIN & VIANEY-LIAUD, 2000
P. hirschi JACKSON & VARRICCHIO, 2010
P. tiantaiensis WANG, ZHAO, WANG & JIANG, 2011
= Elongatoolithus tiantaiensis FANG, WANG & JIANG, 2000
P. Oospecies: Nova VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
P. sp

Oogenus: Protoceratopsidovum MIKHAILOV, 1994
P. sincerum MIKHAILOV, 1994 (Ootype)
P. minimum MIKHAILOV, 1994
P. fluxuosum MIKHAILOV, 1994

Oogenus: Sankofa LOPEZ-MARTINEZ & VICENS, 2012
S. pyrenaica LOPEZ-MARTINEZ & VICENS, 2012 (Type)

Oogenus: Spheruprismatoolithus BRAY, 1999
S. condensus BRAY, 1999 (Ootype)

Gen. sp indet.

MORPHOTYPE: ORNITHOID-PRISMATIC INCERTAE SEDIS

Oogenus: Dispersituberoolithus ZELENITSKY, HILLS & CURRIE, 1996
D. exilis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)

Oogenus: Pseudogeckoolithus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
P. nodosus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997 (Ootype)
Note: Morphotype: Geckonoid?
P. sp

Gen. sp indet.

OOCOHORT: ORNITHISCHIFORMES NOVA

OOSUPERORDER: SAUROPODOMORPHA NOVA

SUBORDER: PLATEOSAURIA TORNIER, 1913

ORDER: PROSAUROPODA HUENE, 1920 (emend)

Family: MASSOSPONDYLIDAE Huene, 1914

Genus: Massospondylus OWEN, 1854
M. sp


TUBOSPHERULITIC STRUCTURAL MORPHOTYPE

oofamily: "MEGALOOLITHIDAE" Zhao, 1979
[Sauropoda]

Note: The first dinosaur eggs remains were discovered in Cretaceous strata of Southern France over a century ago (Pouech..). These eggs, known as megaloolithids, have been assumed to belong to sauropod dinosaurs because they are usually found in strata that are rich in sauropod remains (Erben et al., 1979; Vianey-Liaud et al. 1987; Jain 1989; Sahni et al, 1994). Recent confirmation of this correlation was made with the discovery of South American megaloolithid eggs containing sauropod embryos (Chiappe et al, 1998). Occasional references to typical megaloolithid eggshells as “hadrosaurian’ (Grigorescu et al, 1994, p. 77) are obviously misleading. Additionally, Upper Cretaceous strata in North America are rich hadrosaur remains and contain no trace of megaloolithid eggshell (Zelenitsky, pers. comm.). All positively identified hadrosaurian eggs have a different eggshell structure (see Spheroolithidae).

Oogenus: Cairanoolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
= Dughioolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
C. dughii VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994 (Ootype)
C. roussetensis (VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994) BARCIA & VIANEY-LIAUD, 2001
= Dughioolithus roussetensis VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994

Oogenus: Megaloolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. mammilare VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994 (Ootype)
M. siruguei VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. aurelinesis VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. petralta VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. cylindricus KHOLSA & SAHNI, 1995
= Megaloolithus rahioliensis MOHABEY, 1998
M. jabalpurensis KHOLSA & SAHNI, 1995
= Megaloolithus patagonicus CALVO, ENGLLAND, HEREDIA & SALGADO, 1997
= Megaloolithus matleyi MOHABEY, 1997
M. mohabeyi KHOLSA & SAHNI, 1995
= Megaloolithus phensaniensis MOHABEY, 1998
M. baghensis KHOLSA & SAHNI, 1995
= Megaloolithus pseudomamillare VIANEY-LIAUD, HERISCH, SAHNI & SIGE, 1997
= Megaloolithus balasinorensis MOHABEY, 1998
M. dholiyaensis KHOLSA & SAHNI, 1995
M. padiyalensis KHOLSA & SAHNI, 1995
M. walpurensis KHOLSA & SAHNI, 1995
= Megaloolithus khempurensis MOHABEY, 1998
Note: VIANEY-LIAUD, KHOSLA & GARCIA, 2003 place Megaloolithus walpurensis within the M. khempurensis because of the fragmentary nature of M. walpurensis, but M. walpurensis has priority.
M. trempii MORATALLA, 1998
M. dhoridungriensis MOHABEY, 1998
M. megadermus MOHABEY, 1998
M. microtuberculata GARCIA & VIANEY-LIAUD, 2001
M. maghrebiensis GARCIA, TABUCE, CAPPETTA, MARANDAT, BENTALEB, BENABDALLAH, & VIANEY-LIAUD, 2003
M. nova PANDES I BLAS, 2005
M. sp

Oogenus: Patagoolithus SIMON, 2006
P. salitralensis SIMON, 2006 (Ootype)

Gen. sp indet.

Incertae sedis

Oogenus: Tacuaremborum MONES, 1980
T. oblongum MONES, 1980 (Ootype)

oofamily: FAVEOLOOLITHIDAE Zhao & Ding, 1976 [Sauropoda]

Note: MIKHAILOV, 1997…Only bones of sauropod dinosaurs are known from the Faveoloolithus nest-bearing layers in Ologoy-Ulan-Tsav locality in Mongolia Gobi (Sochava, 1969; S. M. Kurzanov, pers. comm.). The arrangement of the eggs and porosity of the eggshell appear to be similar to those subterranean nesters, such as chelonian.

Oogenus: Faveoloolithus ZHAO & DING, 1976
F. ningxiaensis ZHAO & DING, 1976 (Ootype)
F. sp

Oogenus: Hemifaveoloolithus WANG, ZHAO, WANG & JIANG, 2011
H muyushanensis WANG, ZHAO, WANG & JIANG, 2011 (Ootype)

Oogenus: Parafaveoloolithus ZHANG, 2010
P. microporus ZHANG, 2010 (Ootype)
P. macroporus ZHANG, 2010
P. tiansicunensis ZHANG, 2010
P. guoqingsiensis (FANG, WANG & JIANG, 2000) WANG, ZHAO, WANG & JIANG, 2011
= Dendroolithus guoqingsiensis FANG, WANG & JIANG, 2000
P. pingxiagensis ZOU, WANG & WANG, 2013

Oogenus: Sphaerovum MONES, 1980
S. erbeni MONES, 1980 (Ootype)
S. sp

Oofamily: SIMILIFAVEOLOOLITHIDAE Wang, Zhao,Wang & Jiang, 2011

Oogenus: Simlifaveoloolithus WANG, ZHAO, WANG & JIANG, 2011
S. shuangtangensis (FANG, LU, JIANG & YANG, 2003) WANG, ZHAO, WANG & JIANG, 2011 (Ootype)
= Dendroolithus shuangtangensis FANG, LU, JIANG & YANG, 2003

oofamily: YOUNGOOLITHIDAE Zhang, 2010

Oogenus: Youngoolithus ZHAO 1979
Y. xiaguanensis ZHAO, 1979(Ootype)
Y. xipingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
Y. sp

Gen. sp indet.

Family: Unknown.

Gen. sp indet.

Family: Titanosauridae?

Hypselosaurus priscus?

Gen. sp indet.

oofamily: DENDROOLITHIDAE Zhao and Li, 1988
[Sauropoda? Therizinosauria? Mikhailov, 1997

Note: As per Mikhailov, 1997…Based on the taphonomic settings and reconstructed type of nesting, dendroolithid remains have been attributed tentatively to some ornithopod or sauropod dinosaurs (Mikhailov, 1991). Recently, some privately owned dendroolithid eggs from Henan Province, China displayed at the University Museum of Zoology, Cambridge (the author was kindly allowed to examine shell samples) have been shown to contain well preserved embryos of therizinosaurid dinosaurs at late stages of development (Cohen et al, 1995)…

Oogenus: Dendroolithus ZHAO & LI, 1988
D. wangdianensis ZHAO & LI, 1988 (Ootype)
D. verrucarius MIKHAILOV, 1994
D. microporosus MIKHAILOV, 1994
D. xichuanensis ZHAO & ZHAO, 1998
D. furcatus XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. dendriticus XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. sanlimiaoensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. zhaoyingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. hongzhaiziensis ZHOU, REN, XU & GUAN, 1998
D. tumiaolingensis ZHOU, REN, XU & GUAN, 1998
D. fengguangcunensis FANG, ZHANG, ZHANG, LU, HAN & LI, 2005

Oogenus: Paradendroolithus ZHOU, REN, XU & GUAN, 1998
P. ginglongshanensis ZHOU, REN, XU & GUAN, 1998 (Type)

Oogenus: Placcolithus ZHAO & ZHAO, 1998
P. taohensis ZHAO & ZHAO, 1998 (Ootype)

Gen. sp indet.

oofamily: DICTYOOLITHIDAE Zhao, 1994

Oogenus: Dictyoolithus ZHAO, 1994
D. hongpoensis ZHAO, 1994 (Ootype)

Oogenus: Paradictyoolithus WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
P. zhuangqianensis WANG, ZHAO, WANG, ZHANG, & JIANG, 2013 (Ootype)
P. xiaxishanensis WANG, ZHAO, WANG, ZHANG, & JIANG, 2013

Oogenus: Protodictyoolithus WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
P. neixiangensis (ZHAO, 1994) WANG, ZHAO, WANG, ZHANG, & JIANG, 2013 (Ootype)
= Dictyoolithus neixiangensis ZHAO, 1994
P. jiangi (LIU & ZHAO, 2004) WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
= Dictyoolithus jiangi LIU& ZHAO, 2004

Oogenus: Stromatoolithus ZHAO, YE, LI, ZHAO & YAN, 1991
S. pinglingensis ZHAO, YE, LI, ZHAO & YAN, 1991 (Ootype)
S. gongzhuligensis (WANG, ZAN, JIN & CHEN, 2005) WANG, ZHAO, WANG, ZHANG, & JIANG, 2013
= Dictyoolithus gongzhuligensis WANG, ZAN, JIN & CHEN, 2005

Gen. sp indet.

oofamily: PHACELOOLITHIDAE Zeng& Zhang, 1979

Oogenus: Phaceloolithus ZENG & ZHANG, 1979
P. huanensis ZENG & ZHANG, 1979 (Ootype)

Gen. sp indet.

oofamily: Incertae sedis ?spherulite or spheruprismatic structural type eggshell.

Oogenus: Parvoolithus MIKHAILOV, 1996
P. tortuosus MIKHAILOV, 1996 (Ootype)
Note: Possible beetle castings ?

Gen. sp indet.

oofamily: Incertae sedis

Oogenus: Placoolithus non ZHAO, 1979
P. cf. taohensis ZHAO, 1979 (Ootype)

OOMAGNORDER: ORNITHISCHIA SEELEY, 1888
EGGS WITH DINOSAUROID-SPHERULITIC BASIC SHELL TYPE

Gen. sp indet.

oofamily: SPHEROOLITHIDAE Zhao, 1979
[Ornithopoda: some hadrosaurs]

Note: As per Mikhailov, 1997…Spheroolithus eggs from Laiyang have been attributed to hadrosaurs (Young, 1959, pp. 511, 517); the eggs of S. irenensis were found in close association with Bactrosaurus remains (Zhao and Jiang, 1974, p. 81). Spheroolithid eggs from the Upper Cretaceous Two Medicine Formation of Montana were found in association with hadrosaurian juveniles (Horner and Makela, 1979). Lambeosaurine embryos (Hypacrosaurus stebingeri; Horner and Currie, 1994) were also found in association with large spherical eggs; however, the eggshell morphotype could not be identified as spheroolithid because it is diagenetically modified (Zelenitsky, 1995). This was misinterpreted by Mikhailov. These discoveries have allowed all spheroolithid remains to be attributed to hadrosaurs (Hirsch and Quinn, 1990, Mikhailov et al, 1994). Their abundance, as well as their geographical and stratigraphical distribution, are in accordance with this conclusion…

Oogenus: Paraspheroolithus ZHAO, 1979
Note: (= Spheroolithus ZHAO, 1979 as per MIKHAILOV, 1997)
P. irenensis (CHAO & CHIANG, 1974) ZHAO, 1979 (Ootype)
= Spheroolithus irenensis (CHAO & CHIANG, 1974) MIKHAILOV, 1994
= Oolithes spheroides YOUNG, 1954
= Oolithes irenensis CHAO & CHIANG, 1974
= Spheroolithus sp MIKHAILOV, 1991
P. lamelliformae ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996)
P. shizuiwanensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
P. yangchengensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
P. sanwangbacunensis FANG, ZHANG, ZHANG, LU, HAN & LI, 2005
P. sp

Oogenus: Spheroolithus ZHAO, 1979
S. spheroides (YOUNG, 1954) ZHAO, 1979 (Ootype)
= Oolithes spheroides YOUNG, 1954
S. chiangchiungtingensis (CHAO & CHIANG, 1974) ZHAO, 1979
= Oolithes chiangchiungtigensis CHAO & CHIANG, 1974
S. megadermus (YOUNG, 1959) ZHAO, 1979
= Oolithes megadermus YOUNG, 1959
S. tenuicorticus MIKHAILOV, 1994
S. maiasauroides MIKHAILOV, 1994
S. albertensis ZELENITSKY & HILLS, 1997
S. choteauensis JACKSON & VARRICCHIO, 2010
S. europaeus SELLES, VIA, & GALOBART, 2014
Oosp (unnamed)
= Genus: Maiasaura HORNER & MAKELA, 1979
M. peeblesorum HORNER & MAKELA, 1979
S. sp

Oogenus: Shangyangoolithus ZHANG & XUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996)
= Shanyangoolithus ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996) (sic)
S. shanyangensis ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996) (Ootype)

Oogenus: Shixingoolithus ZHAO, YE, LI, ZHAO, & ZHANG, 1991
S. erbeni ZHAO, YE, LI, ZHAO, & ZHANG, 1991 (Ootype)

Oogenus: Nova XU, ZHAO, LU, DONG & HUANG, 1999

Gen. sp indet.

Incertae sedis

Oogenus: Triprismatoolithus JACKSON & VARRICCHIO, 2010
T. stephensi JACKSON & VARRICCHIO, 2010 (Ootype)

oofamily: OVALOOLITHIDAE Mikhailov, 1991
(?Ornithopoda)

Note: As per Mikhailov, 1997…The histrostructure of the shell and other, superficial features of the egg, are similar to those of the Spheroolithidae. At the Mogoyn-Ulagiyn-Kheates locality (= Tel-Ulan-Ula; Mongolia, Dornogov Aimak), pinkish strata containing numerous ovaloolithid eggs in clutches yield remnants of only lambeosaurine dinosaurs (S. M. Kurzanov, pers. comm.). This prompts the tentative assignment of ovaloolithid remains to ornithopod dinosaurs. Preliminary assignment of embryonic limb bones associated with an ovaloolithid egg (PIN 2970/9) to protoceratopsian dinosaurs (L. I. Hozatcky in Sochen, 1972) has proved to be incorrect (S. M. Kurzanov, see Mikhailov et al., 1994)…

Oogenus: Ovaloolithus ZHAO, 1978
O. chinkangkouensis (CHAO & CHIANG, 1974) ZHAO, 1978 (Ootype)
= Oolithes chinkangkouensis CHAO & CHIANG, 1974
O. laminadermus (CHAO & CHIANG, 1974) ZHAO, 1979
= Oolithes sp YOUNG, 1954
= Oolithes laminadermus (YOUNG, 1954) CHAO & CHIANG, 1974
O. tristriatus ZHAO, 1979
O. mixtistriatus ZHAO, 1979
O. monostriatus ZHAO, 1979
O. dinornithoides MIKHAILOV, 1994
O. sangpingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
O. utahensis BRAY, 1999
O. tenuisus BRAY, 1999
O. sp

Gen. sp indet.

Ornithischia incertae sedis?

HYPSILOPHODONTIDAE

Gen. sp indet.

RHABDODONTIDAE

Gen. sp indet.

DRYOSAURIDAE

Gen. sp indet.

CAMPTOSAURIDAE

Gen. sp indet.

HADROSAURIDAE

Telmatosaurus transsylvanicus

Gen. sp indet.

LAMBEOSAURINAE

Hypacrosaurus stebingeri

PSITTACOSAURIDAE

Gen. sp indet.

PROTOCERATOPIDAE?

Protoceratops

Gen. sp indet.

CERATOPIAN

In general

KENTROSAURINAE

Dacenterurus armatus

OOCLASS: AVES NOVA

Incertae sedis

Oogenus: Plagioolithus IMAI & AZUMA, 2014
P. fukuiensis IMAI & AZUMA, 2014 (Ootype)

Gen. Sp indet.

oofamily: LAEVISOOLITHIDAE Mikhailov, 1991 [AVES, Enantiornithes]
= oofamily: SUBTILIOLITHIDAE Mikhailov, 1991 [AVES]

Note: As per Mikhailov, 1997… These small and thin-shelled eggs with ornithoid eggshell structure were proposed to belong either to birds or small theropod dinosaurs (Mikhailov, 1991). Recently, one of the specimens of laevisoolithid eggshells was found in situ with an associated partial skeleton of an enantiornithid bird Nanantius (Kurockhin, 1996). This allows taxonomic assignment of laevisoolithid eggs to Enantiornithes (Mikhailov, 1996a)

Oogenus: Laevisoolithus MIKHAILOV, 1991
L. sochavai MIKHAILOV, 1991 (Ootype)

Genus: Gobipteryx ELZANOWSKI, 1976
G. minuta ELZANOWSKI, 1976
= Nanantius valifanovi KUROCHKIN, 1996
G. sp

Oogenus: Subtiliolithus MIKHAILOV, 1991
S. microtuberculatus MIKHAILOV, 1991 (Ootype)
S. kachchhensis KHOLSA & SAHNI, 1995

Oogenus: Tipoolithus GARCIA, TABUCE, CAPPETTA, MARANDAT, BENTALEB, BENABDALLAH, & VIANEY-LIAUD, 2003
T. achloujensis GARCIA, TABUCE, CAPPETTA, MARANDAT, BENTALEB, BENABDALLAH, & VIANEY-LIAUD, 2003 (Ootype)

Gen. sp indet.

oofamily: GOBIOOLITHIDAE Mikhailov, 1996 [Aves]

Note: As per Mikhailov, 1997…Well-developed embryos of palaeognathous birds have been found at Khermiyn-Tsav (Elzanowski, 1981; Kurochkin, 1988). Although Elzanowski never stated that these embryos should be assigned to the small bird Gobipteryx minuta, described earlier from deposits of the same age (Elzanowski, 1977), this opinion has occasionally appeared in the literature Recent re-examination of embryonic material (Kurockhin, 1995) has shown that the embryo, which shares some features with Gobipteyx, is not certainly associated with the Gobioolithus minor eggs. Embryos that are certainly from Gobioolithus minor eggs may belong to a different group of birds, apparently Palaeognathae…

Genus: Gobipipus CHATTERJEE, 1994 (noma nudum)
G. elzanowskii CHATTERJEE, 1994 (noma nudum)

Oogenus: Gobioolithus MIKHAILOV, 1996
G. minor MIKHAILOV, 1996 (Ootype)
G. maior MIKHAILOV, 1996

oofamily: ORNITHOOLITHIDAE Fang, in Fang, Zhang, Pang, Li, Han, Xie, Yan, Pang Lu & Cheng, 2007

Oogenus: Protornithoolithus FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG, LÜ & CHENG, 2007
P. tumendongensis FANG, in FANG, ZHANG, PANG, LI, HAN, XIE, YAN, PANG, LÜ & CHENG, 2007 (Ootype)

Morphotype: Ornithoid-ratite

Incertae sedis.

Oogenus: Ageroolithus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
A. fontllongensis VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997 (Ootype)
A. sp

Oogenus: Continuoolithus ZELENITSKY, HILLS & CURRIE, 1996
C. canadensis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably small theropod.
C. sp

Oogenus: Porituberoolithus ZELENITSKY, HILLS & CURRIE, 1996
P. warnerensis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably small theropod.
P. sp

Oogenus: Tristraguloolithus ZELENITSKY, HILLS & CURRIE, 1996
T. cracioides ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably belongs to a bird.

Oogenus: Tubercuoolithus JACKSON & VARRICCHIO, 2010
T. tetonensis JACKSON & VARRICCHIO, 2010 (Ootype) .

Gen. sp indet.

Bibliography

Bibliography for taxa list

_____________________________________________________________________________________

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A

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B

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C

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D

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E

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F

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G

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H

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I

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J

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K

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L

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M

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N

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O

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S

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V

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W

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X

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Y

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Z

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