Species: tetragonus SEELEY, 1895
Etymology:

Holotype: SAM-PK-571b

Locality: Burghersdorp, Albert District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull fragment.

Referred material:

BRINK, 1977

Locality: Kalkfeld, Namibia.

Horizon: Omingonde Formation, Moletno, Etjo beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSO R329: Fragmentary skull.

 

BRINK, 1986

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP F3774: Skull.

 

Locality: Leeuspruit, Rouxville District, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone A, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP F4658: Small skull.

 

Locality: Winnaarsbaken, Burgersdorp, Albert District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

PBI F3511: Skull.

 

DAMIANI, 2008

Locality: Farm Cuylerville (annex of the farm Prospect 152), Queenstown District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone, subzone B.

Age: Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

Number: Not given:

 

GRINE, 1977

Locality: Wonderboom, South Africa.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

SAM 571b: Skull.

 

HENDRICKX, ABDALA & CHOINIERE, 2019

Locality: Burgersdorp District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

NHMUK PV R2574, R2575, SAM-PK-3426, K175, K180, K183, K9968:

 

Locality: Winnaars Baken, Burgherdorp District, Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP/1/3511, NHUK PV R4092, R9216, SAM-PK-8015, 11265:

 

Locality: Aliwal North, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

NHMUK PV R3308, R3724, SAM-PK-4002, 5877, 6216, 6218, 6219:

 

Locality: Queenstown, Eastern Cape Provine, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

SAM-PK-?K4660,?K4661:

 

Locality: Hofmeyr, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B or C, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

SAM-PK-k5266:

 

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP/1/1195, 3758, 505, UMCZ T.438, T.441, T.445, T454, T.826, T.971:

 

Locality: Bethel/Slookrall, Rouxville District, Free State, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone A, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP/1/4529, 4647:

 

Locality: Gladde Grond 530 locality, Rouxville District, Free State, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone A, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

PB/1/4669:

 

Locality: Betjieskraal 36 locality, Rouxville District, Free State, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone A, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP/1/4677:

 

Locality: Norwood Farm, Rouxville District, Free State, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone C, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP/1/5541:

 

Locality: Avilion Farm, Rouxville District, Free State, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone C, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP/1/5542:

 

Locality: Unknown, Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

NHMUK PV R3736, K5223, K5716, K8971, K9969:

 

Locality: Beardmore Glacier Region, Antarctica.

Horizon: Fremouw Formation.

Biostratigraphy:

Age: Early Traissic.

Material:

AMNH FR 24421:

KEYSER, 1973

Locality: Etjo Mountian, Namibia, South West Africa.

Horizon: Omingonde Formation, Moletno, Etjo beds.

Biostratigraphy:

Age: Early Triassic.

Material:

GSN R327: Fragmentary skull, and fragmentary skeleton.

GSN R335, R321: Fragmentary lower jaws.

GSN RK3:

 

MARTINELLI, FUENTE & ABDALA, 2009

Locality: In the vicinity of the Puesto Viejo Farm House, about 40 km southwest of San Rafael, Mendoza Province, Argentina.

Horizon: Rio Seco de la Quebrada Formation, Puesto Viejo Group.

Biostraigraphy:

Age: Ansian Stage, Lower Mid Triassic Epoch, Lower Middle Triassic.

Material:

MNSR-Pv 357: Skull fragments of a single individual including partial lateral right portion of the skull (snout and temporal regions), a fragment of the left snout with dentition, left zygomatic arch, small portion of skull roof with a fragment of the occipital plate and the lateral wall of the braincase, and fused dentaries with partial dentition lacking postdentary bones.

 

PEECOOK, STEYER, TABOR & SMITH, 2017

Locality: Lower Ntawere (Drysdall & Kitching's Locality 16) north of Sitwe, upper Luangwa Valley, Eastern Province, northeastern Zambia.

Horizon: Ntawere Formation.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

BP/1/3638: A well-preserved skull an dmandibles in articulation.

= Cynochampsa laniaria OWEN, 1860
Etymology:
= Diademodon? laniaria (OWEN, 1860)
= Cyclogomphodon laniaria (OWEN, 1860) BROOM, 1919
.
Holotype: BMNH 36229

Locality: Rhenosterberg, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Fragmentary skull and dentary.

Referred material:

OWEN, 1876

BMNH A. 43 (old): 4 thoracic vertebrae and a left coracoid.

BMNH S. A. 44 (Old): Proximal and distal portions of a left humerus.

 

= Diademodon brachytiara SEELEY, 1894
Etymology:
= Diademodon brachytious (sic?)

Holotype: SAM-PK-571a

Locality: Wonderboom, Burgersdorp District, Eastern Cape Province, South Africa.

Horizon:

Biostratigraphy: Subzone B, Cynognathus Assemblage Zone.

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: 2 isoalted upper postcaines.

 

= Diademodon entomophonus SEELEY, 1895
Etymology:

Holotype: BMNH R 3765/4092

Locality: Winnaarsbaken, Burgersdorp District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Imperfect skull.

Referred Material:

GRINE, 1977

Locality: Nooitgedacht, Burgersdorp District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

NM C2705, C2707A, C2707B, C2707C, C2707D, C2707E, C2707F, C2707G, C2707H, C2707I, C2707J: Skulls.

 

= Gomphognathus browni SEELEY, 1895
= Diademodon browni (SEELEY, 1895)
Etymology:

Holotype: NHMUK PV (old BMNH) R3304

Locality: Aliwal North, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull fragment.

Referred material:

BRINK & KITCHING

Locality: Winnaarsbaken in the Burgersdorp District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

Bernard Price Institute 225: Nearly complete skull.

 

BROILI & SCHRODER, 1935

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

MUNICH 1934 VIII 14: Skull.

 

GRINE, 1981

BP.FN 3511, 3769, 3773, 3772: Skulls.

 

KITCHING

Locality: Across the Donga near the Grootdam boundary fences, Burgersdorp district, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

Number: Not given:

 

 

 

= Gomphognathus kannemeyeri SEELEY, 1895
Etymology: Dr. Daniel Rossouw Kannemeyer (1843-1925), the South African fossil collector.
= Diademodon kannemeyeri (SEELEY, 1895)

Holotype: Albany museum 438, 458 (457?)

Locality: Burgersdorp, Albert District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull fragment.

 

= Gomphognathus mastacus SEELEY, 1895
Etymology:
= Diademodon mastacus (SEELEY, 1895)

Holotype: NHMUK PV R3303 (SAM 1332?)

Locality: Aliwal North, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull fragment.

Referred material:

Locality: South Africa.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

DMSW R51: Fragmentary skull.

 

BROOM

Locality: Farm Luiper Kop, Burgersdorp District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

M314/F2522: Sagittal half of a large skull.

HENDRICKZ, ABDALA & CHOINIERE, 2019

UMCZ T.433:

BRINK, 1955

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

DMS R152, R153: Fragmentary skulls.

531, DMS R205, R204: Fragmentary skeletons.

 

BROILI & SCHRODER, 1935

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

MUNICH 1934 VIII 15: Skull.

MUNICH 1934 VIII 16: Skull and dentary.

MUNICH 1934 VIII 20: Skull missing tip of snout and dentary.

 

BROOM, 1904

Locality: Aliwal North, Eastern Cape Province, South Africa..

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

Number: Not given: Fragmentary dentary and fragmentary skeleton.

 

GRINE, 1981

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BPL.FN 3756:

SAM 1332, USNM 23352, UMCZ (old CAMB) T434, MUNICH 1934 VIII 30: Skulls.

 

KITCHING

Locality: Due north of Burgersdorp on the Odendallstroom road lies the farm Luiperdkop, Burgersdorp District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

Number: Not given: A good sagittal half of a skull.

HENDRICKZ, ABDALA & CHOINIERE, 2019

BP/1/2522:

= Gomphognathus polyphagus SEELEY, 1895
Etymology:
= Diademodon polyphagus (SEELEY, 1895)

Holotype: NHMUK PV (old BMNH) R2576-7

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Imperfect skull.

Referred material:

GRINE, 1981

NHMUK PV (old BMNH) R2578, SAM K.5222: Skulls.

 

= Gomphognathus dimorphodon SEELEY, 1908
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

 

= Gomphognathus minor BROOM, 1911
Etymology: Greek, minor, "minor, small."

Holotype: NHMUK PF (old BMNH R) 3587/3588

Locality: Winnaars Baken, Burgherdorp District, Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

NHMUK PF R 3587: Skull without anterior lobe of snout.

NHMUK PF R 3588: Skull.

 

= Cyclogomphodon platyrhinus BROOM, 1913
Etymology:
= Diademodon platyrhinus (BROOM, 1913)

Holotype: AMNH FR 5519

Locality: Winnaars Baken, Burgherdorp District, Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull and lower Jaws.

Referred material:

AMNH FR 5518:

BRINK, 1986

Locality: Matyantya, Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP M319: Anterior half of skull.

 

Locality: Luiperdskop, Burgersdorp, Albert District, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

BP M320: Anterior half of skull.

 

= Octagomphus woodi BROOM, 1919
Etymology:
= Diademodon woodi (BROOM, 1919)

Holotype: Albany Museum 3753

Locality: Steynsburg, Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Imperfect skull.

 

= Protacmon brachyrhinus WATSON, 1920
Etymology:

Holotype: CAMB T 462

Locality: Essex, Prince Albert Distict, Western Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Nearly complete skull and dentary.

 

= Gomphognathus grossarthi BROILI & SCHRODER, 1935
Etymology:
= Diademodon grossarthi (BROILI & SCHRODER, 1935)

Holotype: MUNICH 1934 VIII 17

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull.

Referred material:

BROOM

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

M359/:

BP/1/3754: Skull.

 

BROILI & SCHRODER, 1935

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

Munch Samml Nr. 1934 VIII 17: Skull.

 

= Gomphognathus haughtoni BROILI & SCHRODER, 1935
Etymology: In honor of Sidney Henry Haughton (1888-1982), British Paleontologist.
= Diademodon haughtoni (BROILI & SCHRODER, 1935)

Holotype: MUNICH 1934 VIII 19

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull.

Referred material:

BROOM

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

M360/:

BP/1/3757: Preorbital half of a very large skull.

 

= Gomphognathus broomi BROILI & SCHRODER, 1935
Etymology: Robert Broom (1866-1951), Scottish-born paleontologist who worked in South Africa.
= Diademodon broomi (BROILI & SCHRODER, 1935)

Holotype: MUNICH 1934 VIII 18

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull missing snout.

Referred material:

Locality: Farm Cragievar about a mile south of Burgersdorp, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

M361/F3772: Immature skull, without lower jaw, rather complete with some skeletal bones.

M362/F3773: Immature skull, without lower jaw.

M362/F3769: Immature skull and lower jaw.

M364/F3771: Immature skull and lower jaw, with some additional skeletal
bones.

 

BRINK & KITCHING

Locality: Immediately north of the river running in a westerly direction north of the town Lady Frere, on the slope of a koppie, Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material:

RC 313: Fragmentary Skull.

 

= Sysphinctostoma smithi BROILI & SCHRODER, 1936
Etymology:

Holotype: BSP 1936 II 8

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull of a juvenile and lower jaw.

 

= Sysphinctostoma gracilis BROOM, 1950
Etymology: Latin, gracilis, "slender."

Holotype: RC 113

Locality: Father Ruebsamen for Lumku, Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull and fragmentary skeleton.

 

= Protacmon reubsameni BROOM, 1950
Etymology:

Holotype: RC 112

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Fragmentary skull.

 

= Gomphodontoides megalops BRINK & KITCHING, 1951
Etymology:

Holotype: Bernard Price Institute 222

Locality: Winnaarsbaken in the Burgersdorp district, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Fragmentary skull of a juvenile.

 

= Diademodon? laticeps BRINK, 1955
Etymology:

Holotype: DMS R131

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Skull.

Referred material:

UMCZ T 436: Skull.

 

= Diademodon parringtoni BRINK, 1955
Etymology:

Holotype: DMS R159

Locality: Lady Frere District, Transkei, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Fragmentary skull and dentary.

 

= Diademodon rhodesiensis BRINK, 1955
Etymology:

Holotype: BP/1/3639

Locality: Luangwa Valley, Northern Zambia, East Africa.

Horizon: Ntawere Formation.

Biostratigraphy:

Age: Early Triassic.

Material: Skull missing tip of snout.

 

= Cragievarus kitchingi BRINK, 1965
Etymology:

Holotype: BP/1/ 3776.

Locality: Below and west of the Wonderboomspruit railway bridge on the farm Cragievar, 5 miles south of Burgersdorp, Eastern Cape Province, South Africa.

Horizon: Burgersdorp Formation, Tarkastad Subgroup.

Biostratigraphy: Subzone B, Cynognathus zone (Kannemeyeria-Diademodon zone, KEYSER, 1979, SACS, 1980, Kannemeyeria zone, KEYSER & SMITH, 1977-77).

Age: Spathian Stage, uppermost Scythian Epoch to Anisian Stage, lowermost Mid Triassic Epoch, Early to Middle Triassic.

Material: Fragmentary skull.

Referred material:

HENDRICKZ, ABDALA & CHOINIERE, 2019

BP/1/3769, 3772-3773: