Genus: Anhanguera CAMPOS & KELLNER 1985
Etymology: Tupi culture, indigenous tribe of Brazil, Anhanga, 'devil', and nera, 'what was, old'.
= Anghanguera WELLNHOFER, 1988 (sic)
= Anhanguera CAMPOS vide LEONARDI, 1984 (nomen nudum)
= Tropeognathus WELLNHOFER, 1987 (partim)
Etymology: Greek, tropis, 'keel', and Greek, gnathos, 'jaw'; named for the keeled jaws.

Species: blittersdorffi CAMPOS & KELLNER 1985
Etymology: In honor of Mr. Rainer Alexander von Blittersdorff.

Holotype: MN 4805-V (originally in the private Collection Desiree, from Mr. Rainer Alexander von Blittersdorff)

Locality: The exact site is unknown, however this material came from calcareous sediments of the Chapada do Araripe, probably of a flank in Ceara State, NE Brazil.

Horizon: Santana Formation, Araripe Group.

Biostratigraphy:

Age: Aptian-Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Skull lacking all the teeth except three.

Referred material:

KELLNER & TOMIDA, 2000

Universidade Estadual do Rio Janeiro (n. 40 Pz-DBAV-UERJ): Partial skull and mandible.


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Species: santanae (WELLNHOFER, 1985) KELLNER, 1990
Etymology:
= Araripesaurus santanae WELLNHOFER, 1985

Holotype: BSP 1982 I 90

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Skull missing distal end, dentary missing distal and proximal ends, ulna and radius, 2 right proximal carpals, 3 right distal carpals and proximal portion of right metacarpal IV.

Referred material:

= Anhanguera cf. santanae APPLEGATE, 1999
Note: Was Pteranodon or Nyctosaurus in CABRAL-PERDOMO & APPLEGATE, 1994
= Genus: Nova? APPLEGATE & CABRAL-PEERDOMO, 1994

Locality: Tlayua in Tepexi de Rodriguez, Puebla State, Central Mexico.

Horizon: Tlayua Formation.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Number: Not given: A humerus 2 scapulocoracoids.

Other remains are fragments, and complete phalanages for the wing finger, a fragment of a pteroid bone, and a small femur.

 

WELLNHOFER, 1988

Locality: Barra do Jardim, Chapada do Araripe, Ceara State, North-east Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

AMNH 22555: Nearly complete skull, the articular end of the right mandible, the complete vertebral series including the sacrals but without the caudals, several ribs, both scapulae and coracoids, the proximal parts of both humeri, both carpal regions with fragments of radius, ulna and metacarpals. The pelvis is almost complete and preserved three-dimensionally along with the proximal ends of both femora and a few phalanges of the foot.


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Species: araripensis (WELLNHOFER, 1985) KELLNER & TOMIDA, 2000
Etymology: In reference to Chapade do Araripe.
= Santanadactylus araripensis WELLNHOFER, 1985
= Coloborhynchus araripensis (WELLNHOFER, 1985) VELDMEIJER, MEIJER & SIGNORE, 2006

Holotype: BSP 1982 I 89

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Mid-section of skull, anterior portion of dentary, both ceratobranchialia 1 of the hyoid apparatus, right humerus, distal end of right ulna, distal end of right radius, right proximal carpal, right
distal carpal, right distal lateral carpal fragment, proximal end of right metacarpal III, proximal portion of right metacarpal IV, sesamoids of the right carpal area.
Note: Left ramus with a necroses pit, right mandibular ramus has bony overgrowth (BENNETT, 2003).

Modified from Wellnhofer, 1985.

Referred material:

= cf. Anhanguera araripensis KELLNER & TOMIDA, 2000

MN 4735-V: A complete skull.

 

BSP 1982 I 92: Left humerus, missing the deltrapectorial crest.

 

VELDMEIJER, MEIJER & SIGNORE, 2006

St. Gallen SAO 16494: Skull.

 

WELLNHOFER 1991

Locality: Barra do Jardim, Chapada do Araripe, Ceara State, North-east Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

Bayerische Staatssammlung fur Palaontologie und historische Geologie, Munchen, BSP 1987 I 66: Wing finger metacarpal.

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Species: robustus (WELLNHOFER, 1987) KELLNER & CAMPOS, 1988
Etymology:
= Tropeognathus robustus WELLNHOFER, 1987
= Coloborhynchus robustus (WELLNHOFER, 1987) VELDMEIJER, 1998

Holotype: BSP 1987 I 47

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Lower dentary and teeth.

Paratype: SMNK 2303 PAL: Anterior part of the upper and lower jaws.

Referred material:

VELDMEIJER, 1998

Leiden: Skull and lower jaw?

 

UNWIN, FREY, MARTILL, CLARKE & RIESS, 1996

Locality: Chapade do Araripe, Ceara State, Northeastern Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

SMNK 1133PAL: Fragmentary skull with mandible, and parts of the postcranial skeleton.
Note: With a 6 m wingspan.

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Species: piscator KELLNER & TOMIDA, 2000
= Coloborhynchus piscator (KELLNER & TOMIDA, 2000) UNWIN, 2001
Etymology: The species name piscator comes from the Latin and means fisherman, in allusion to the assumed fishing activity of this pterosaur.
= Anhanguera sp KELLNER, & TOMIDA, 1993

Holotype: NSM-PV 19892

Locality: Araripe Basin, and was possibly collected in the proximity of Santana do Cariri, Ceara State, Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Partial skeleton, consisting of skull, mandible, part of the vertebral column (atlas and axis, 6 cervicals, 5 dorsals, 4 sacrals, 11 caudals), 2 right ribs, incomplete sternum, left scapula and coracoid, proximal articulation of the right coracoid, left humerus, left radius and ulna, distal articulation of the right radius and ulna, both complete carpal regions, part of both pteroids, complete left McIII and wing metacarpal (McIV) and distal parts of the left McI and McII, proximal part of the right McIII and the wing metacarpal (McIV), proximal articulation of the left first wing phalanx (F1D4), proximal part of the right second wing phalanx (F2D4), incomplete right third wing phalanx (F3D4), incomplete right fourth wing phalanx (F4D4), digits I to III of the left manus, complete left femur, incomplete right femur, incomplete tibias, and incomplete pes.

Referred material:

A complete cast is deposited at the Museu de Ciencias da Terra of the Departamento Nacional da Producao Mineral (MCT 1535-R) and a second at the Museu Nacional/UFRJ (MN 5023-V), Rio de Janeiro.

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Species: spielbergi (VELDMEIJER, 2003) RODRIGUES & KELLNER, 2008
= Coloborhynchus spielbergi VELDMEIJER, 2003a
Etymology: In honor of Steven Spielberg, the director of the three Jurassic Park movies in which dinosaurs and pterosaurs were animated.

Holotype: RGM 401 880

Locality: Unknown, Chapada do Araripe, Ceara State, North-east Brazil.

Horizon: Romualdo Member, Santana Formation, Araripe Group.

Biostratigraphy:

Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Skull, mandible, 7th and 8th cervical vertebrae, notarium, left and right scapulocoracoid, sternum, left humerus, left ulna, left radius, left proximal syncarpal, a 3rd phalanx, 3 claws, 7 dorsals, 5 pieces of ribs, pelvis, both femora and tibiae and fragments of ceratobranchialia.

 

Referred material:

= cf. Coloborhynchus spielbergi VELDMEIJER, 2002

SMNS 5413: Right ulna.