Genus: Nicrosaurus O. FRAAS, 1866
Etymology:
= Lophorhinus MEHL, 1915 (sic)
= Lophosaurus MEHL, 1915 vide HAY, 1902/FITSINGER, 1843 (sic)
= Lophoprosopus MEHL, 1915
Etymology:
= Microsaurus ROMER, 1956/DeJEAN, 1833 (sic)

Species: kappffi (MEYER, 1861) FRAAS, 1866
Etymology:
= Belodon kappffi MEYER, 1861
= Lophoprosopus kappffi (MEYER, 1861) MEHL, 1915
= Phytosaurus kappffii (MEYER, 1861) McGREGOR, 1906

Paralectotype: SMNS 54708

Locality: Probably from Heslacher Wand, Stuttgart-Heslach (Von Lowensein, Stuttgart, Wildenau), Baden-Wurttemberg State, Germany.

Horizon: Lowenstein Formation (old Stubensandstein, Middle Keuper).

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material: Anterior half of a left premaxilla.

Lectotype: SMNS 4060 and SMNS uncat. no. 15: Badly corroded and damaged snout fragment.

Referred material:

HAVLIK, AIGLSTORFER, ATFY, & UHL, 2013

Locality: Federal highway construction site B29 - Ortsumgehung Schwäbisch Gmünd "Gmünder Einhorn Tunnel" in Schwäbisch Gmünd, Baden-Württemberg State, Germany.

Horizon: Lower Löwenstein Formation.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

GPIT/RE/7341: Long bone fragment.

GPIT/RE/7335, 7337, 7339, 7342: 4 teeth.

 

HUNGERBUHLER, 2000

Locality: Gaisburg, Near Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Middle Stubensandstein, Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

SMNS 5727: Skull.

SMNS 23078: prenarial portion of a snout of a large individual with about 30 well-preserved teeth.

 

Locality: From former quarries near the villages of Heslach and Kaltental, in the vicinity of Stuttgart, Baden-Wurttemberg State, Southern Germany.

Horizon: Middle Stubensandstein, Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

SMNS 4378: Almost complete skull.

SMNS 4379: Complete and almost undistorted skull.

SMNS 5725: Preorbital part of a large skull.

SMNS 5726: Complete skull.

SMNS 4380: Complete mandible.

SMNS 5730: Right mandibular ramus with 26 teeth, five of with are artificial.

 

LYDEKKER, 1888

Locality: Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

BMNH 39477: Cast of the cranium.

BMNH 42743: The Skull lacking most of the posterior parts behind the supratemporal fenestrae.

BMNH 38037: Posterior portion of skull.

BMNH 38039: Fragmentary palato-maxilla.

BMNH 38040: Fragmentary premaxilla.

BMNH 38041: Fragmentary skull.

BMNH 38042: Fragment of left temporal region.

BMNH 38043: Fragment of left temporal region.

BMNH 38044: Right partial.

BMNH 37994: Fragmentary skull.

BMNH 38036: Complete mandible with 64 teeth preserved.

BMNH 42744: Complete mandible with a total of 19 teeth preserved.

BMNH 38045: Fragmentary right dentary with 3 teeth.

BMNH 38046: Fragmentary left mandibular symphysis of a juvenile.

BMNH 38047: Right splenial.

BMNH 38048: Fragmentary left spleinal.

BMNH 38049: Fragmentary right splenial.

BMNH 38050: Fragmentary right splenial.

BMNH 38052: Fragmentary right dentary.

BMNH 38068: Teeth.

BMNH 38060-61: 2 bones not improbably belonging to the hyoid apparatus.

BMNH 38069: 2 cervical vertebrae.

BMNH 38070: Fragmentary 6th dorsal vertebra.

BMNH 38071-79: Imperfect dorsal vertebrae.

BMNH 38080: 4 caudal centra.

BMNH 38081: A chevron.

BMNH 38082: The arch and part of the centrum of the 8th (?) dorsal vertebra.

BMNH 42746: Fragmentary dorsal vertebra.

BMNH 38091: 3 fragmentary ribs.

BMNH 38047: An abdominal rib.

BMNH 38067: Fragmentary right scapula of a juvenile.

BMNH 38066: Left coracoid.

BMNH 38055: Fragmentary right humerus.

BMNH 38056: Left humerus.

BMNH 38063: Left ilium.

BMNH 38064: Fragmentary right ilium.

BMNH 38065: Pubis.

BMNH 38054: Right femur.

BMNH 38059: Left tibia.

BMNH 38083: Dorsal scute.

BMNH 38084: Fragmentary right dorsal scute.

BMNH 38085: Fragmentary dorsal scute.

BMNH 38086: Fragment right dorsal scute.

BMNH 38087: A small dorsal scute.

BMNH 38088: Dorsal scute.

BMNH 38089: 3 dorsal scutes.

BMNH 38090: Imperfect dorsal scutes.

BMNH 42748: Fragmentary dorsal scute.

BMNH 42748a: Fragmentary dorsal scute.

 

MEYER, 1861

Locality: Heslach, near Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Stubensandstein, Middle Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

SMS? Elements of skulls, and skeletal elements, and dorsal scutes.

Note: As per HUNT, 1994

Lectotype: SMNS partial rostrum (MEYER, 1861, Pl. 30).

Paralectotype: SMNS rostral tip (MEYER, 1861, pl. 31, fig. 9-12.)

Referred material:

SMNS: 3 skulls on display.

SMNS 5729: Partial skull.

 

HUENE, 1911

Locality: Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Upper Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

No. 5726: Skull.

No. 5735: Fragmentary skull.

 

HUENE 1908, GALTON 1984, 1985

Locality: Heslach, near Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Middle Stubensandstein, Middle Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

SMNS 5715: Right ilium.

 

HUENE, 1922

Locality: Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

Nr. 4391: Right femur.

Nr. 12671: Dorsal vertebrae.

Nr.? Left ischium, right humerus, left ulna, right tibia, left fibula, metatarsal V, IV, III, II, I.

 

Locality: Pfaffehnofen, Baden-Wurttemberg State, Germany.

Horizon: Stubensandstein, Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

Nr.? Scapula and coracoid, interclavical, right ilium.

Nr. 12986: Femur, ilium ischium and pubis.


_____________________________________________________________________________________

Species: meyeri HUNGERBUHLER & HUNT, 2000
Etymology: In honor of Christian Hermann von Meyer (1801-1869), for his pioneering contributions to our knowledge of Triassic vertebrates that includes the first descriptions of phytosaur skulls referable to the new genus.
= Belodon plieningeri MEYER, 1860 (partim)
= Phytosaurus plieningeri (MEYER, 1842) LYDEKKER, 1888
= Phytosaurus kappffii (MEYER, 1842) ABEL, 1923 (partim)

Holotype: SMNS 13240

Locality: Quarry “Weißser Steinbruch”, Pfaffenhofen, northern Baden-Wurttemberg State, Germany.

Horizon: Middle Stubensandstein, subunit sc 1, “werkstein” beds, Stubensandstein, Middle Keuper.

Biostratigraphy:

Age: Middle Norian Stage, Late Triassic Epoch, Late Triassic.

Material: Skull in two pieces.

Referred material:

SMNS 12593/2: Skull lacking the snout.

BMNH 38038: Right mandibular ramus.

GPIT 261/011: Right mandibular ramus.


Locality: Kayh, Schleswig-Holstein State, Germany.

Horizon: Middle Stubensandstein, subunit sc 1, “werkstein” beds, Stubensandstein, Middle Keuper.

Biostratigraphy:

Age: Middle Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

SMNS 4059: partial skull.

BMNH 42745: Complete skull.

BMNH 38039: Snout fragment.

BPT 2070.001: Partial skull.

NHM (BM[NH]) R. 3582: Skull.

 


= Phytosaurus plieningeri LYDEKKER, 1888

Locality: Stuttgart, Baden-Wurttemberg State, Germany.

Horizon: Keuper.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

BMNH 42745: Nearly complete skull.

BMNH 38038: Right ramus and teeth.
Note: Nicrosaurus sp nova HUNGERBUHLER, 1999