Genus: Edaphosaurus COPE, 1882
Etymology:
= Brachycnemius WILLISTON, 1911
Etymology:
= Naosaurus COPE, 1886
Etymology:

Species: cruciger (COPE, 1878) ROMER & PRICE, 1940
Etymology:
= Dimetrodon cruciger COPE, 1878
= Naosaurus cruciger (COPE, 1878)

Holotype: Not identifiable.

Locality: Northwestern Archer County, Texas.

Horizon: Belle Plains Formation, Whichita Group or Clyde Formation, Clear Fork Group.

Horizon: Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group or Lower to middle Waggoner Ranch Formation (formerly Clyde Formation, upper Wichita Group) Upper Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B or Land-vertebrate faunachron lower to middle C.

Age: Upper Wolfcampanian Stage, Upper Rotliegendes Epoch, Early Permian or Lower Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Lectotype: AMNH 4060

Locality: From near the mouth of Beaver Creek, Northwest of Dundee, Archer County, Texas.

Horizon: Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: 29 continuos vertebrae from axis to base of tail.

After ROMER & PRICE, 1940, Skeleton from AM 4060, skull from WM 658.

After ROMER & PRICE, 1940

Referred material:

AMNH 4039 (partim): Fragmentary spines and vertebrae.

 


Locality: From the valley of the Little Wichita River, Archer County, Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

AMNH 4003: Several fragmentary centra, several fragmentary spines.

 


Locality: Locality 35, Ohio River drainage, Pittsburgh-West Virginia.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous) or Early Permian.

Material:

No. 8540: Spine fragment, scraps of vertebrae, ribs and a partial tooth plate.

 


Locality: Locality 25, Ohio River drainage, Pittsburgh-West Virginia.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous) or Early Permian.

Material:

Number: Not given: Spine fragments.

 

 

Locality: Locality 28, Ohio River drainage, Pittsburgh-West Virginia.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous) or Early Permian.

Material:

Number: Not given: Spine and rib fragment.

 

 

Locality: Locality 37, Ohio River drainage, Pittsburgh-West Virginia.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous) or Early Permian.

Material:

Number: Not given: Spine and rib fragments and a clavicle.

 

CASE, 1915

Locality: Brier Creek Bone-bed, about 12 miles south of Dundee, a little east of the road between Dundee and Archer City, in Archer County, Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

3333: Left humerus.

 

CASE, 1918

Locality: Brier Creek Bone-bed, about 12 miles south of Dundee, a little east of the road between Dundee and Archer City, in Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

University of Michigan: Fragmentary skeleton.

 


LYDEKKER, 1890

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

BMNH R750: Fragment of maxilla that may belong to Dimetrodon.

BMNH R564: 2 portions of a neural spine.

BMNH R564a: Part of the neural arch and spine of a dorsal vertebrae.

 


OLSON, 1967

Locality: Southwest Taylor Site, Secs, 3, 4, T5S, R11W, Cotton County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 4-7-S22: Spine.

OUSM 4-40-11: Vertebrae, limb bones and ribs.

 


Locality: McCann Quarry Site, the quarry side on Deer Creek, 8 miles west and 4 miles south of Blackwell (about 9 miles southwest in a direct line) and 2.5 miles northeast of Eddy, In Kay County, NE 1/4, sec. 16 and SW 1/4, sec. 9, T26N, R2W, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 4-35-S2: Spine.

OUSM 4-35-S4: Complete spine and fragmentary spine.

 

ROMER & PRICE, 1940

Locality: Near Slippery Creek, south of Dundee, Archer County, Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

FMNH UC 658: Skull.

 


Locality: Near the mouth of Beaver Creek in southwestern Wichita County, Texas.

Horizon: Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

AMNH 4072: Spine.

 


Locality: Tit Mountain, just northeast of Dundee, Archer County, Texas.

Horizon: Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ: Fibula.

 


Locality: From the western end of Baylor County, North of Maybelle, Texas.

Horizon: Lower to middle Waggoner Ranch Formation (formerly Clyde Formation, upper Wichita Group) Upper Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle C.

Age: Lower Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

FMNH WM 658: Skull.

 


Locality: Elmo, Dickenson County, Kansas.

Horizon: Probably Clyde Formation equivalent, Lower to middle Waggoner Ranch Formation (formerly Clyde Formation, upper Wichita Group) Upper Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle C.

Age: Lower Leonardian Stage, Lower Rotliegendes Epoch, Early Permian..

Material:

Number: Not given: Fragments.

 

WHIPPLE & CASE, 1930

Locality: Portland, Jackson County, West Virginia.

Horizon: Dunkard Group.

Biostratigraphy:

Age: Wolfcampanian stage, upper lower to lower middle Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Spines, centra and ribs.

 


= Naosaurus microdus COPE, 1881
Etymology:
= Edaphosaurus microdus (COPE, 1881) COPE, 1884

Holotype: AMNH 4014

Locality: From near the mouth of Beaver Creek, Northwest of Dundee, Archer County, Texas.

Horizon: Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: Fragmentary spines, a few vertebrae, and 2 fragments of the dentigerous plate of the pterygoids.

Referred material:

AMNH 4060: A nearly perfect vertebral column from the 3rd cervical to the 2nd or 3rd caudal.

 

 

 

_____________________________________________________________________________________

Species: pogonias COPE, 1882 (Type)
Etymology:
= Naosaurus pogonias COPE, 1882

Holotype: AMNH 4009

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Fragmentary skull and lower jaw and imperfect axis.

After ROMER & PRICE, 1940

After ROMER & PRICE, 1940, Backbone taken without addition from WM 163, skull, scapulocoracoid, and humerus from WM 239, hind leg WM from 1092, Clavicle from AM 4037, Pelvis from AM 4002.

From CASE, 1907.

Referred material:

AMNH 4037: Spine material and clavicle.

FMNH WM 186: Scapulocoracoid.

FMNH WM 16: 2 pelvic girdles, not associated.

AMNH 4038: Fragments of vertebrae and spines

FMNH WM 1099: Tibia.

 


Locality: Moonshine Creek, Baylor Creek, Texas.

Horizon: Clear Fork beds.

Biostratigraphy:

Age: Early Permian.

Material:

FMNH UC 625: Skull.

 


Locality: Not given.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

University of Michigan 3333: Fragmentary skull, scapula, pelvis, limb bones.

 

ROMER & PRICE, 1940

Locality: North of Seymour, county seat of Baylor County, Grey's Creek Dransi northward into the Big Wichita, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

FMNH WM 163: Vertebral column.

MCZ 1117: Partial scapulocoracoid.

NM 10461: Several vertebrae and spines.

 


Locality: Hog Creek rise west of Table Top Mountian as a southern tributary of the Big Wichita, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

From CASE, 1907

AMNH 4015 (1348): Presacral column.

 


Locality: Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

FMNH WM 239: Skull, scapulocoracoids, humerus and incomplete column.

After ROMER & PRICE, 1940

Locality: Unknown.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

AMNH 4030: Fragmentary spine and a few centra.

AMNH 4017: Spine.

 

 

ZIDEK, JOHNSON, MAY & CLABORN, 2004

Locality: OMNH V 173, Northeast Frederic site of Simpson, (1976, 1979), Tillman County, Oklahoma.

Horizon: Garber Sandstone, Upper part of the Sumner Group, possibly equivalent to the Lower Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

Number: Not given:

 

= Naosaurus claviger COPE, 1886
Etymology:

Holotype: AMNH 4002

Locality: Coffee Creek near the Big Wichita River, in Vernon County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: 26 vertebrae, fragmentary spines, pelvis and several ribs.

 

Referred material:

BROILI, 1904

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

Number: Not given: Scapula and coracoid.

 

LYDEKKER, 1890

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

BMNH R564b: Distal half of the neural spine of a dorsal vertebra.

BMNH R564c: Fragment of spine.

 

REISZ, 1986

Locality: Craddock Bonebed, north of Seymour, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material:

FMNH UC 163: A vertebral column.

 


= Brachycnemius dolischomerus WILLISTON, 1911
Etymology:

Holotype: FMNH WM 1092

Locality: Hog Creek rises west of Table Top Mountains as a southern tributary of the Big Wichita, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Femur, tibia and fibula.

_____________________________________________________________________________________

Species: novomexicanus WILLISTON & CASE, 1913
Etymology:

Holotype: FMNH UC 674

Locality: Arroyo de Agua, in the Vicinity of Arroyo de Agua, Rio Arriba County, New Mexico.

Horizon: Cutler/Abo Formation.

Biostratigraphy:

Age: Wolfcampian Stage, Lower Rotliegendes Epoch, Early Permian.

Material: Fragmentary skull, 15 articulated vertebrae with the skull, fragmentary scapulae, right humerus and radius.

Referred material:

MODESTO & REISZ, 1992

CM 34910: Partial tibia, two metatarsals, phalanges, and several anterior caudal vertebrae.

 


Locality: El Cobre Canyon, north of Abiuin, Rio Arriba County, North-central New Mexico.

Horizon: Abo/Cutler Formation.

Biostratigraphy:

Age: Wolfcampian, Lower Rotliegendes Epoch, Early Permian.

Material:

UCLA VP 1641: Several neural spine fragments.

 


Locality: Cottonwood Canyon, Southern New Mexico.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous).

Material:

UCLA VP 1719: 4 closely associated posterior dorsal vertebrae and rib fragments.

 

ROWLAND, LUCAS, KUES & ESTEP, 1997

Locality: New Mexico.

Horizon: Madera Formation, Red Tanks Member.

Biostratigraphy:

Age: Virgilian, Gzelian Epoch, Late Pennsylvanian (Late Late Carboniferous).

Material:

Number: Not given: Neural spines.

 

VAUGHN, 1969

Locality: NW 1/4, sec. 6, T15S, R11E, Domingo Canyon, Tularosa Locality, Otero County, New Mexico.

Horizon: Abo Formation.

Biostratigraphy:

Age: Wolfcampian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Partial neural spines.

_____________________________________________________________________________________

Species: boanerges ROMER & PRICE, 1940
Etymology:

Holotype: MCZ 1531 (partim)

Locality: Geraldine Bonebed, Geraldine, south of the Geraldine School, approximately 13 km northwest of Archer City, Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material: Jaw rami.

After ROMER & PRICE, 1940

Compsite from ROMER & PRICE, 1940

Note: As per MODESTO, 1995, ‘This number originally contained ‘the remains of about six individuals’ from Geraldine, with the type jaws forming part of a mounted MCZ specimen. In the early 1980’s all specimens except the type jaws were reassigned to MCZ numbers 4309-4324.’

Referred material:

MCZ 1680: A partial skeleton with an obliquely compressed skull.

MCZ 1762: A partial skeleton with fragmentary skull and mandibles.

MCZ 1764: A fragmentary skull.

MCZ 4309: A partial skull with mandibles (formerly part of MCZ 1531).

ROM 37760: A fragmentary left maxilla.

Note: As per MODESTO, 1995, ‘This is not an exhaustive list of Edaphosaurus boanerges specimens riposted at the MCZ, ROM, or elsewhere, and included only those specimens examined here.’

 


OLSON, 1967

Locality: Waurika Site 1, 6 miles west of Waurika in SW 1/4, NW 1/4, sec. 36. T4S, R9W, Jefferson County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 4-7-S15: 5 fragments of small individuals.

OUSM 4-7-S20: Spine fragment.

 


Locality: Southeast Paoli site, NW 1/4, Sec. 19, T4N, R1E, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 4-7-S11: Fragment of spine.

 

ROMER, 1952

Locality: Localities 8 and 12, Ohio River drainage, Pittsburgh-West Virginia.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous) or Early Permian.

Material:

Number: Not givin: Spine fragments.

 


Locality: Localities 8 and 12, Ohio River drainage, Pittsburgh-West Virginia.

Horizon:

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous) or Early Permian.

Material:

No. 8579: An ilium, partial humerus, metapodial and spine fragment.

 

ROMER & PRICE, 1940

Locality: Brier Creek, in western Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

FMNH WM 703, UM 3333: Fragmentary skeletons.

UM 3446: Braincase.

FMNH WM 819: Maxilla.

UM 3758: Axis.

FMNH WM 56: Interclavicle.

FMNH WM 534: Clavicle.

UM 1163: Scapulocoracoid and humerus.

FMNH WM 456: Humerus.

UM 1166: Radius and ulna.

UM 1165, 1164: Femora.

FMNH WM 703: Tibia.

AMNH 4769, FMNH WM 563: Vertebrae and spines.

UM : Fibula.

 


Locality: Table Branch, in the western headwaters of the South Fork in Southern Archer County, Texas.

Horizon: Upper Archer City Formation (formerly Putnam Formation, Wichita Group), upper Bowie Group.

Biostratigraphy: Land-vertebrate faunachron upper A.

Age: Wolfcampanian stage, Lower Rotliegendes Epoch, Early Permian.

Material:

MCZ 1372: Posterior trunk, sacral and proximal caudal vertebrae with much of spines.

MCZ 1370: Lower jaw.

 

 


Locality: Rattlesnake Canyon, southeast of Black Flat in Northwestern Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

MCZ 1286: The greater part of the presacral column and spines.

 


Locality: Exposures west of Anarene, Archer County, in the slopes east of the South Fork of the Little Wichita River, Texas.

Horizon: Upper Archer City Formation (formerly Putnam Formation, Wichita Group), upper Bowie Group.

Biostratigraphy: Land-vertebrate faunachron upper A.

Age: Wolfcampanian stage, Lower Rotliegendes Epoch, Early Permian.

Material:

MCZ 1629: Jaw.

 

 


Locality: Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group. or Petrolia Formation (formerly Belle Plains Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower to upper B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

AMNH 4003, 4080: Vertebrae and spines.

UM 3021: Femur.

 


SANDER, 1989

Locality: Archer City Bonebed 2, Archer County, Texas.

Horizon: Upper Archer City Formation (formerly Putnam Formation, Wichita Group), upper Bowie Group.

Biostratigraphy: Land-vertebrate faunachron upper A.

Age: Wolfcampanian stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Unprepared skeleton at TMM.

 


Locality: Archer City Bonebed 3, Archer County, Texas.
Horizon: Upper Archer City Formation (formerly Putnam Formation, Wichita Group), upper Bowie Group.

Biostratigraphy: Land-vertebrate faunachron upper A.

Age: Wolfcampanian stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Neural spines.

JOHNSON, 2012

SMU:

 

 

Locality: Coprolite Bonebed, Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: Neural spines.

 

 

Locality: Loftin Bonebed, Archer County, Texas.

Horizon: Admiral Formation, Wichita Group.

Biostratigraphy:

Age: Leonardian stage, Upper Rotilegendes Epoch, Early Permian.

Material:

Number: Not given: Neural spines.

 


Locality: Rattlesnake Canyon Bonebed 2, Archer County, Texas.

Horizon: Nocona Formation (formerly lower to lower upper Admiral Formation, middle Wichita Group), Lower Wichita Group.

Biostratigraphy: Land-vertebrate faunachron lower B.

Age: Upper Wolfcampanian stage, Upper Rotliegendes Epoch, Early Permian.

Material:

Number: Not given: ?Vertebrae.


_____________________________________________________________________________________

Species: colohistion BERMAN, 1979
Etymology:

Holotype: CM 23513

Locality: Road cut on Interstate Highway 70 about 1/2 mile east of Elm Grove, West Virginia.

Horizon: Limestone "B", Pittsburgh Formation, Monogohela Group.

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous).

Material: 14 essentially complete presacral vertebrae believed to include cervicals 6 and 7 and dorsals 8-19, disarticulated but closely associated with the partial vertebral column are 4 or 5 intercentra, 2 incomplete vertebrae and numerous neural spine fragments near both ends of the column, and numerous ribs.