PROVINCEIA DO BEIRA LITORAL
TO THE NORTHEAST OF SERRA DE BOA VIAGEM
VIZO (= VISO)
D’ARAZEDE
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Rosasia soutoi
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: VARANIDAE Gray, 1827
Gen. sp indet.
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
POMBAL
VALE DE PORTHINHEIRO
UNNAMED UNIT
OXFORDIAN- KIMMERIDGIAN STAGE
LOWER-MIDDLE MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
non Megalosaurus, M? pombali
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
2 KM NE OF POMBAL
? UPPER KIMMERIDGIAN STAGE
LOWER-MIDDLE MALM EPOCH TO LOWER TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
Subfamily: KENTROSAURINAE nova
Dacentrurus armatus
= Omosaurus lennieri
= Dacentrurus lennieri
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
POMBEL
LOWER KIMMERIDGIAN STAGE
LOWER-MIDDLE MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
non Megalosaurus M? insignis
= Erectops insignis
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
DISTRICT OF COIMBRA
TAVEIRO VILLAGE
TAVEIRO FORMATION
LATE CAMPANIAN OR EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
MICRORDER: MARGINOCEPHALIA SERENO, 1986
ORDER: PACHYCEPHALOSAURIA MARYANSKA & OSMOLSKA, 1974
SUBORDER: DOMOCEPHALINIA SERENO, 1986
INFRAORDER: PACHYCEPHALOSAURIA (MARYANSKA & OSMOLSKA, 1974) COOPER, 1985
Incertae sedis
*Taveirosaurus costai
= cf. Goyocephale sp
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
DISTRICT OF COIMBRA
TAVEIRO
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: TRIBOSPHENIDA (McKENNA, 1975) McKENNA & BELL, 1997
SUPERCOHORT: THERIA (PARKER & HASWELL, 1987) McKENNA & BELL, 1997
COHORT: MARSUPIALIA (ILLIGER, 1811) McKENNA & BELL, 1997
Incertae sedis
Gen. sp indet.
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
TAVEIRO
UPPER CAMPANIAN-MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
TAVERIO
EMPRESA DE CERAMICA VOUGA
LATE CAMPANIAN OR LOWER MAASTRICHTIAN STATE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
*Rosasia soutoi
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
Gen. sp indet.
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: VARANIDAE Gray, 1827
Gen. sp indet.
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
TAVEIRO
MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
UPPERMOST GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Rosasia soutoi
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
DISTRICT OF COIMBRA
TAVEIRO VILLAGE
LATE CAMPANIAN OR EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
non Megalosaurus M. pannoniensis
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
DISTRICT OF COIMBRA
TAVEIRO VILLAGE
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet. A.
Gen. sp indet B.
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
Gen. sp indet.
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPODA PAUL, 2002
SUBORDER: COELURIA MARSH, 1881
Family: COELURIDAE Marsh, 1881
Gen. sp indet.
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
*Euronychodon portucalensis
= Paronychodon lacustrisGen. sp indet.
Family: TROODONTIDAE Gilmore, 1924
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: HOMALOSAUROPIA JANENSCH, 1929
PARVORDER: DIPLODOCIMORPHA CALVO & SALGADO, 1995
Superfamily: DIPLODOCOIDEA Marsh, 1884 (sensu Upchurch, 1995)
Family: DIPLODOCIDAE Marsh, 1884
Gen. sp indet.
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF BEIRA LITORAL
VIZO (= VISO)
UPPER CAMPANIAN-MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: VARANIDAE Gray, 1827
Gen. sp indet.
SUPERORDER: SERPENTES LINNAEUS, 1759
ORDER: OPHIDIA BRONGNIART, 1800
SUBORDER: ALETHINOPHIDIA NOPCSCA, 1923
Superfamily: BOOIDEA Gray, 1825
Family: BOOIDAE
Gen sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
Incertae sedis
Gen. sp indet.
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
Incerate sedis
Gen. sp. indet
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
(Small theropod)
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
non Megalosaurus M. pannoniensis
CLASS: AVES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF DO BEIRA LITORAL
ABOUT 25 WNW OF COIMBRA
VIZO (= VISO)
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF BEIRA LITORAL
VIZO (= VISO)
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
ORDER: NEOTETANURAE SERENO, WILSON, LARSSON, DUTHEIL & SUES, 1994
SUBORDER: CARNOSAURIA HUENE, 1920
Superfamily: Incertae sedis
Family: MEGALOSAURIDAE Huxley, 1889
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
PORTUGAL
PROVINCE OF BEIRA LITORAL
VIZO (= VISO)
RAILWAY CUTTING NEAR VIZO (= VISO)
LATE CAMPANIAN OR EARLY MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
Incertae sedis
Gen. sp indet.
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
Incertae sedis
Gen sp indet.
(Small theropod)
_____________________________________________________________________________________