Genus: Diplocaulus COPE, 1877
Etymology:
= ?Permoplatyops CASE, 1946
Etymology:
= Peronedon OLSON, 1970
Etymology:
= Platyops WILLISTON, 1918 non TWELVETREES, 1880

Species: salamandroides COPE, 1877
Etymology:

Syntypes: FMNH UC 6513-16

Locality: Horseshoe bend of Vermillion River, Danville, Vermillion County, Illinois.

Horizon: "Clepsydrops shales", Middle Mcleansboro Formation, Conemaough Group.

Biostratigraphy:

Age: Late Pennsylvanian (Late Carboniferous).

Material: 4 presacral vertebrae.

Referred material:

PATON, 1975

Locality: Oakwood, Vermilion County, Illinois

Horizon:

Biostratigraphy:

Age: Asselian Stage, Rotliegendes Epoch, Early Early Permian.

Material:

RSM: 1 vertebra.

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Species: magnicornis COPE, 1882
Etymology:

Lectotype: AMNH 4539

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Posterior border of skull and 4 anterior vertebrae.

Referred material:

AM 4466, 4467, 4468, 4469, 4472, 4473, 4478, 4494, 4498, 4501, 4504, 4509, 4512, 4527, 4528, 4530, 4538, 4544, 4589, 4597, 4752, CNHM UC 207. 221, 229, 636, 637, 1013, 1317, 1650, 1652, 1653, 1654, 1656, 1657, 1658, 1660, 1663, USNM 17884: Fragmentary skulls.

AM 4523a, 4523b, 4752, 4485, 4589, 4597, UC 206, 1660, 224, 222, 1663, 1658, 223: Fragmentary skulls of larvae.

AM 4511, AM 4491, UC 1656, 1657, 1652, 410, 1650, P12689: Fragmentary skulls of juveniles.

WILLISTON,

BROILI, 1904

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

Fragmentary skull, and skeletal elements.

COPE, 1895

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material: Nearly complete skull.

 

DOUTHITT, 1917

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material: Skulls and skeletons.

 

 

 

OLSON, 1967

Locality: West Grandfield Site, SE 1/4, sec. 4, T4S, R15W and SE 1/4, sec. 32, T3S, W., Tilman County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-4-S12: Lower Jaw.

 

Locality: Northwest Edmond Site, 7 miles northwest of Edmond, NW 1/4, NE 1/4, sec. 25, T15N, R4W, Logan County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-1-S17: 2 skull fragments.

OSUM 2-10-S18: Skull fragment.

 

= Diplocaulus cf magnicornis JOHNSON, 1994

Locality: Orlando Site, 2 miles northeast of Orlando, SE 1/4, sec. 31, T20N, R1W, 2 miles northeast of Orlando in Noble County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given:

 

OLSON, 1967

Locality: Deep Red Run Locality, Sec. 21, T4S, R12W, Cotton County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-4-S11: Posterior part of jaw.

OUSM 2-7-S8: Vertebrae.

 


Locality: Perry Site 6, SE 1/4, SE 1/4, sec. 26, T23N, R2W, Noble County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-40-S9: Fragments of skull, jaw, and vertebrae.

 

Locality: Orlando Site, 2 miles northeast of Orlando, SE 1/4, sec. 31, T20N, R1W, 2 miles northeast of Orlando in Noble County, Oklahoma.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

CNHM UR 1026: Jaw, and fragments.

CNHM UR 1027: Vertebrae, and partial jaw.

CNHM UR 1028: Parts of small individuals.

CNHM UR 1029: Fragments of skull and vertebrae.

CNHM UR 1030: about 100 miscellaneous pieces of many individuals.

 

Locality: Mc Cann Quarry Site, the quarry is on Deer Creek, 8 miles west and 4 miles south of Blackwell (about 9 miles southwest in a direct line) and 2.5 miles northeast of Eddy, NE 1/4, sec. 16 and SW 1/4, sec. 9, T26N, R2W, in Kay County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-1-S10: Two-thirds of a skull.

 

Locality: Lucien Site 1, NE 1/4, NE 1/4, sec. 11, T21N, R3W, about 4 miles north of Lucien, Noble County, Oklahoma.

Horizon: Willington Formation (Garber Formation), Summer Group, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Middle Leonardian stage, upper Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-0-S16: Column and fragmentary skull.

OUSM 2-40-S2: Skull fragment, atlas, and vertebrae.

OUSM 2-40-S3: Many skull fragments, girdles and vertebrae.

OUSM 3-4-S1: Lower jaw.

 

Locality: Bean Farm Site, about 10 miles northeast of Edmond in Logan County, SW 1/4 sec 24, T15N, R2W, Oklahoma.

Horizon: Lower Willington Formation (Lower Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Middle Leonardian stage, upper Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-0-S11: Well-preserved skull and skeleton.

 

Locality: South Covington Site, Garfield County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM unnumbered

 

Locality: Pond Creek Site, NE 1/4, NW 1/4, sec. 14, T25N, R5W, about 5 miles east of the village of Pond Creek, Grant County, Oklahoma.

Horizon: Willington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Middle Leonardian stage, upper Rotliegendes Epoch, Early Permian.

Material:

OUSM 2-0-S19, 2-1-S20, 2-1-S29, 2-1-S36: Skull fragments and parts.

OUSM 2-7-S2: 2 vertebrae.

OUSM 2-40-S1: Skull fragments, vertebrae, etc. of several individuals.

OUSM 2-40-S8: Miscellaneous skull fragments.

 

Locality: South Covington Site, 6 miles south of Covington, Garfield County, Oklahoma.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material:

OUSM 2-0-S17: Skull fragments.

 

OLSON, 1970

Locality: Bed 16, about 9 miles northeast of Perry, Logan County, Oklahoma, SE 1/4, SE 1/4, sec. 26, T23N, R2W.

Horizon: Wellington Formation (Garber Formation), Summer Group, Arbuckle limestone, (Probably equivalent to the Arroyo Formation of Texas).

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

Number: Not given:

 

= Diplocaulus limbatus COPE, 1895
Etymology:

Holotype: AM 4471

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Posterior border of skull and 4 anterior vertebrae.

 

= Diplocaulus pusillus BROILI, 1904
Etymology:

Holotype:

Locality: Texas.

Horizon:

Biostratigraphy:

Age: Early Permian.

Material: Fragmentary skull of a juvenile.

Referred material:

3 more fragmentary skulls of juveniles.

 

= Platyops parvus WILLISTON, 1918
Etymology:
= Permoplatyops parvus (WILLISTON, 1918) CASE, 1946

Holotype: CNHM UC 206

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Posterior border of skull and 4 anterior vertebrae.

 

= Diplocaulus primigenius MEHL, 1921
Etymology:

Holotype: CNHM UC 564

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Posterior border of skull and 4 anterior vertebrae.


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Species: copei BROILI, 1904 (nomen vanum)
Etymology:

Syntypes: BSM 1901 XV 44, 45, 47

Locality: Seymour, Willbarger County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: 3 imperfect specimens.

Note: Diplocaulus copei was rejected as indeterminate by CASE 1911: 22, and OLSON, 1951: 93. D. copei is a nomen vanum restricted to the syntypes only.

Referred material:

4 more fragmentary skulls and fragmentary skeletons.

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Species: brevirostris OLSON, 1951
Etymology:

Holotype: AMNH 4470

Locality: Coffee Creek, Baylor County, Texas.

Horizon: Arroyo Formation, Lower Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower D.

Age: Middle Leonardian stage, upper Rotliegendes, Early Permian.

Material: Skull, vertebrae, and parts of appendicular skeleton.

Paratypes:

FMNH UR 1648, 1655, 1661: Fragmentary skulls.
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Species: recurvatus OLSON, 1951
Etymology:

Holotype: FMNH UR19

Locality: Locality KD, approximately 7 miles north of Vera, Knox County Texas.

Horizon: Upper Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material: Skull with part of left anterior-lateral margin missing.

Paratypes:

FMNH UC 1664: Partial skull with much of left horn missing.

FMNH UR 21: Impression of right horn.

Referred material:

Locality: Locality KH, approximately 7 miles north of Vera, Knox County, Texas.

Horizon: Upper Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

FMNH UR 17: Large skull with anterior part missing.

FMNH UR 18: Part of mid-section of large skull.

FMNH UR 22: Impression of much of right horn.

 

Locality: Locality KE, approximately 7 miles north of Vera, Knox County, Texas.

Horizon: Upper Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

FMNH UC 1666: Skull.

 


Locality: Locality KA, approximately 7 miles north of Vera, Knox County, Texas.

Horizon: Upper Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

FMNH UR 20: Fragmentary skull.

FMNH UC 1665: Skull.

 

OLSON, 1972

Locality: "Diplocaulus site", Locality FA, Foard County, Texas.

Horizon: Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

FMNH UR 167: Fragmentary skull, jaws, a series of vertebrae and part of the girdles and limb bones.

FMNH UR 168: Fragmentary skull.

FMNH UR 169: Vertebrae.

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Species: primus (OLSON, 1970) MILNAR, 1996
= Peronedon primus OLSON, 1970
Etymology:

Holotype: FMNH UR 1234

Locality: Southeast of Norman, SW 1/4, NW 1/4, sec. 13, T8N, R2W, Cleveland County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material: Skull, jaws, 10 vertebrae including atlas, ribs, part of shoulder girdle and part of right and left forelimbs.

Referred material:

FMNH UR 1235: Fragments of 2 small individuals on single block of matrix; includes partial skulls and lower jaws, badly damaged, 2 series of vertebrae and associated ribs.

FMNH UR 1236: Large individual, about twice the size of others, consisting of a crushed skull and 2 vertebrae.

FMNH UR 1237: Fragment of skull and lower jaw.

FMNH UR 1238: Skull.

 

DALY, 1973

Locality: South Grandfiled, southern Tillman County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

FM UR 2293: Partial skull.

FM UR 2294: Skull fragments and a few loose vertebrae.

 

HAGLUND, 1977

Locality: South Granfield, Tillman County, Oklahoma.

Horizon: Hennessey Formation, (Arroyo equivalent), Low Clear Fork Group.

Biostratigraphy:

Age: Leonardian Stage, Lower Rotliegendes Epoch, Early Permian.

Material:

UCLA VP3323: Section of vertebral column.

UCLA VP 2345: Right half of skull and four vertebrae.

 

Locality: Vale Site 1919 OLSON and BOLLES 1975, Texas.

Horizon: Vale Formation, Middle Clear Fork Group.

Biostratigraphy: Land-vertebrate faunachron lower to middle D.

Age: Upper Leonardian Stage,Upper Rotliegendes Epoch, Early Permian.

Material:

UCLA VP3247: Partially crushed skull.

UCLA VP3246: Skull.

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Species: parvus OLSON, 1972
Etymology:

Holotype: UCLA VP 3015

Locality: Site BC-1 (Olson 1965), SW 1/4, SW 1/4, sec. 6, T17N, R10W, 2 miles east of Hitchcock, Blaine County, Oklahoma.

Horizon: Chickasha Formation.

Biostratigraphy:

Age: Guadalupian Stage, Lower Zechstein Epoch, Early Late Permian.

Material: Fragmentary skull, vertebrae, ribs, shoulder girdle, humerus, radius and ulna.
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Species: minimus DUTUIT, 1988
Etymology:

Holotype: MHHN ARG 501

Locality: Argana, Atlas occidental, South, Morocco.

Horizon: Argana Formation.

Biostratigraphy:

Age: Autunain, Asselian Stage, Lower Rotliegendes Epoch, Early Permian.

Material: Fragmentary skull and cervical vertebrae.

Referred material:

Top, Type, ARG 501, Bottom,. ARG 500.

MHHN ARG 500: Fragmentary skull.

MHHN ARG 502: Fragmentary skull.


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Note: Diplocaulus has been found in burrows (SCHULTZE, 1985)