Ichnogenus: Pteraichnus STOKES, 1957
Etymology: Greek, Ptero-, pteron, "wing" and Greek, ichnus, "trace," referring
a track or trace fossil.
Ichnospecies: saltwashensis STOKES, 1957
Etymology: In referrnce to the Salt Wash Sandstones, where the specimens were
found.
Holotype: University of Utah.
Locality: Northwest flank of Carrizzo Mountains, Apache Canyon, Apache County, Arizona; unsurveyed territory about 16.5 miles southwest of four corners.
Horizon: Salt Wash Member, Morrison Formation.
Biostratigraphy: Zone 2.
Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.
Material: A trackway consisting of 9 pairs of consecutive footprints on 4 slabs.
Referred material:
BILBEY, MICKELSON, HALL, KIRKLAND,MADSEN, BLACKSHEAR & TODD, 2005
Locality: Flaming George tracksite, Flaming George Reservoir, Uintah County, northeastern Utah.
Horizon: Uppermost Stump Formation.
Biostratigraphy:
Age: Early Late Jurassic (just below the Salt Water Wash Member of the Morrison Formation).
Material:
In situ: Manus and pes prints, trackways.
LOGUE, 1977, 1996
Locality: Near Alcova, Alcova Reservoir, Natron County, Wyoming.
Horizon: Sundance Formation.
Biostratigraphy:
Age: Oxfordian Stage, Lower Malm Epoch, Late Jurassic.
Material:
Werner Wildlife Museum: Manus, pes and body prints.
Werner Wildlife Museum: Another slab has adult and juvenile prints.
LOCKLEY & WRIGHT, 2003
UW 12360-12372, TATE 0049.1-49.12: Swimming trackways.
MEYERS & BREITHAUPT, 2014
Locality: Seminoe Reservoir Tracksite, Carbon County, Wyoming.
Horizon: Windy Hill Sandstone Member, upper Sundance Formation.
Biostratigraphy:
Age: Late Oxfordian-Early Kimmeridgian Stage, Lower Malm Epoch, Late Jurassic.
Material:
Blk 1 TW1-F, Blk 1 TW2-a, -b, -A, -B, -C, Blk2NWT1-a to e, Blk2NWT21-A to -C, BLK6Tw2-J, Blk6-L, Blk6-M, Blk6TW2-N, Blk6-P, Blk6Tw3-Q to S, BLK6-T, -U, -V, Blk6Tw3-x, Blk7WTw1-a, -b, -B, Blk7WT2-a, Blk7WT2-B, Blk7W-a to -e, Blk7WA to G, Blk7W-h, -i, Blk8W-a to c, Blk8W-a, Blk9W-a, Blk0Wa, Blk10E-a, -A, -B, Blk 11E-a, Blk12E-a to F, Blk13E-a, -B, Blk 14E-A, Blk 15SETw-a, Blk 15Etw-b, Blk16Etw1-A, -B, -C, Blk16E-D, Blk 17Etw-a, Blk 17Etw-A, Blk 17ETW-B, Blk 18W-a, Blk 19NW-A, Blk20S-a, Blk21NW-A, Lr1-a, -b, LR2-a, -B, Lr3-a, Blk2NWTw1-D to -F, Blk2NWTw1-a to -c, Blk3NWTw1-B, -C, Blk4SWTw-a, -B, Blk5SW-a, -A, Blk6Tw1-a to -e, Blk6Tw1-A to -e, Blk6Tw2-g, Blk6-H, ?Blk6-k, Blk6Tw2-J, Blk6Tw2-J., Blk6-L, -M, Blk6TW2-N, Blk6Tw2-O, Blk6-P, Blk6Tw3-O to -S, Blk6-T to -V, Blk6Tw3-x, Blk&WT21-a, -b, Tlk7WT21-B, Blk7Wtw2-a, Blk7W7WTw2-A, Blk7W-a to -e, Blk7WA, Blk7W-F, -G, Blk7W-h, -i, Blk8W-a to -c, LR3-B, -c, LR3-C, LR4a to c, ?LR4-D, LR50a, LR5-A, LR7-a, -b, LR7-A, -B, LR8-a to -h, LR8-F, -H, LR9-a, -b, LR9-A, -B, LR9-d, LR9-D, LR9-g to -n, Blk100-A to -d, Blk100-D, Blk101-A, Blk102-a, Blk102-A: 250 pes and manus tracks.
_____________________________________________________________________________________
Ichnospecies: stokesi LOCKLEY, LOGUE, MORATALLA, HUNT, SCHULTZ & ROBINSON,
1995
Etymology: In honor of William Lee Stokes, who first named Pteraichnus and
attributed it to a pterosaurian trackmaker.
Holotype: Tate Mineral Museum, Casper Wyoming, UW 12368
Locality: Alcova Lake, Alcova Reservoir, Natron County, Wyoming.
Horizon: Sundance Formation.
Biostratigraphy:
Age: Oxfordian Stage, Lower Malm Epoch, Late Jurassic.
Material: 4 partial manus pes sets.
Referred material:
UW 12363-67, 12399-12372: Trackways.
University of Wyoming Geological Museum, Laramie, Wyoming.
= Pteraichnus cf. stokesi PASCUAL-ARRIBAS, & HERNANDEZ-MEDRANO, 2016
Locality: La Muela, Soria Province, Spain.
Horizon: Huerteles Formation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
MUE-A-1: Pes.
MUE-A-2: Manus.
MUE-A-3: Pes.
MUE-A-4: Manus.
MUE-A-7: Pes.
MUE-A-10: Manus.
MUE-A-11: Pes.
MUE-B-1: Manus.
MUE-B-2: Pes.
MUE-B-3: Pes.
MUE-B-4: Pes.
MUE-B-5: Pes.
MUE-B-6: Manus.
MUE-C-1: Pes.
MUE-C-2: Manus.
MUE-C-3: Manus.
MUE-C-4: Manus.
MUE-C-5: Manu.
MUE-C-13: Pes.
MUE-D-1: Manus.
MUE-D-2: Manus.
MUE-D-3: Manus.
MUE-D-4: Manus.
MUE-D-5: Pes.
MUE-D-8: Manus.
MUE-E-1: Manus.
MUE-E-2: Manus.
MUE-E-4: Manus.
MUE-E-9: Manus.
MUE-E-10: Manus.
MUE-E-11: Manus.
MUE-E-13: Manus.
MUE-E-15: Manus.
MUE-E-23: Manus.
MUE-2-24: Manus.
MUE-F-1: Pes.
MUE-F-3: Manus.
MUE-F-6: Manus.
MUE-F-8: Pes.
MUE-F-9: Manus.
MUE-G-1: Manus.
MUE-G-2: Manus.
MUE-H-1: Pes.
MUE-H-2: Manus.
MUE-H-3: Pes.
MUE-H-6: Manus.
MUE-H-7: Pes.
MUE-H-8: Pes.
MUE-H-9: Manus.
MUE-H-19: Manus.
MUE-I-1: Manus.
MUE-I-2: Pes.
MUE-I-5: Manus.
MUE-J-1: Manus.
MUE-J-4: Manus.
MUE-K-1: Manus.
MUE-K-2: Manus.
MUE-L-1: Manus.
MUE-F-5: Manus.
_____________________________________________________________________________________
Ichnospecies: palacieisaenzi PASCUAL
ARRIBAS & SANZ PEREZ, 2000 emend BILLON-BRUYAT & MAZIN, 2003
Etymology: Etymology: In honor of Pedro Palacios and Clemente Saenz, two geologist from Soria, Spain.
= Pteraichnus palaciei-saenzi PASCUAL ARRIBAS & SANZ PEREZ, 2000
Holotype: LTR-R2
Locality: “Los tormos”, Santa Cruz de Yaguas, Soria Province, Spain.
Horizon: Sandy shales and sandstones of sub-unit IV-B of the Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material: Manus.
Paratypes:
LTR-R3: Trackway.
VDV-R3-7: Manus.
VDV-R3-6M: Manus.
DE PIE, VDV-R4-1P: Pes.
Topotypes:
LTR-R1, LTR-R4, LTR-5: Trackways.
Referred material:
Numerous tracks, some form a trackway from Los Tormos, Santa Cruse de Yanguas, and Barranco de Valdelavilla, Valdelavilla, Soria Province, Spain.
PASCUAL-ARRIBAS, CANUDO, PEREZ, HERNANDEZ-MEDRANO, CASTANERA, & BARCO, 2015
Locality: Serrantes I, 2 km west of the locality of Villar del Rio, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: Valles de Valdelalosa lies westwards of the abandoned village of Valdelavilla, San Pedro Manrique, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: La Solaniza, in the vicinity of the locality of Vizmanos, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: La Golondrina, near La Golondrina Hill, to the west of Huerteles, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: Valdegen I, in the ravine of the same name, to the south of the locality of Villar del Rio, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: Valdecantos lies 350 m north of the locality of Valdecantos, at the bottom of a ravine, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: Camino de la MMI, at the starting point of the MM pathway, which links the stream of Miraflores with the SO-630 road, roughly 750 m southwards of the locality of Cerbon, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
Locality: Barranco Hondo, 800 m south of the Piqueras tunnel, Almarza, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
In situ: Trackways.
_____________________________________________________________________________________
Ichnospecies: cidacoi FUENTES VIDARTE, 2001 (nomen
dubium)
Etymology: In reference to the Cidacos River, of Spain, where the specimen was found.
Holotype:
Locality: La Fuente Mingo site, Valloria, Oncala Range, Soria Province, lies on the right bank of the Cidacos River, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Lower-Middle Berriansian Stage, Lowermost Neocomian Subepoch, Lowermost Early Cretaceous Epoch, Early Cretaceous.
Material: 250 tracks.
_____________________________________________________________________________________
Ichnospecies: vetustior MEIJIDE FUENTES,
2001 (nomen dubium)
Etymology: Latin, vetustior, "older."
Holotype:
Locality: Valdelalosa I Palaeontological site, in the Oncala Range, one side of the road that goes from the village of San Pedro Manrique to the village of Fuentes de Magana, Soria Province, Spain.
Horizon: Huerteles Ivb alloformaton, Oncala Group.
Biostratigraphy:
Age: Middle-Upper Berriasian Stage, Lowermost Neocomian Subepoch, Lowermost Early Cretacoeus Epoch, Early Cretaceous.
Material: 44 manus and some pes prints.
_____________________________________________________________________________________
Ichnospecies: parvus MEIJIDE
CALVO, MEIJIDE FUENTES, FUNTES VIDARTE & MEIJIDE
FUNTES, 2001
Etymology: Latin, parvus, "little, small."
Holotype: MNS 2003/92/2
Locality: Valloria outcrops, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lowermost Neocomian Subepoch, Lowermost Early Cretacoeus Epoch, Early Cretaceous.
Material: Manus and pes print.
_____________________________________________________________________________________
Ichnospecies: manueli MEIJIDE CALVO, 2001 (nomen
dubium)
Etymology: In honor of Manuel, who discovered the site where the specimens were found.
Holotype:
Locality: Serrantes ravine, within the Vilar del Rio municipal area on the left bank of Cidacos River, Oncala Range, Soria Province, Spain.
Horizon: Huerteles Alloformation, Cameros Weald.
Biostatigraphy:
Age: Lower-Middle Tithonian-Berriasian Stage, Lowermost Neocomian Subepoch, Lowermost Early Cretaceous Epoch, Early Cretaceous.
Material: Trachway.
_____________________________________________________________________________________
Ichnogenus: yanguoxiaensis Peng, Du, Li, & Bai, 2004
Etymology: In reference to Yanguoxia, where the tracks were found.
Holotype?: CUG-PT1-11
Locality: Site 1, Dinosaur tracksite, Yanguoxia, Yougjing County, Gansu Province, China.
Horizon: Hekou Group.
Biostratigraphy:
Age: Early Cretaceous.
Material: Track.
Paratype? : CUG-PT2: Track.
_____________________________________________________________________________________
Icnhongenus: longipodus FUNTES VIDARTE, MEIJIDE CALVO, MEIJIDE FUNTES, & MEIJIDE
FUNTES, 2004
Etymology: Latin, longus, ‘long’, and Latin, peds, ‘foot’.
Holotype: MNS 2003/92/5
Locality: Barranco de Serrantes outcrop, Soria Province, Spain.
Horizon: Huerteles Alloformation.
Biostratigraphy:
Age: Berriasian Stage, Lower Neocomian Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material: 4 pes and 4 manus prints.
_____________________________________________________________________________________
Ichnospecies: koreanensis LEE, LEE,
LÜ & KOBAYASHI, 2008
Etymology: Named after Korea which yielded the holotype.
Holotype: KIAGAM VP 200401
Locality: An abandoned quarry next to Hadong power plant, Hadong county, South Gyeongsan Province, South Korea.
Horizon: Hsandong Formation, Sindong Group.
Biostratigraphy:
Age: Late Aptian or Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.
Material: Natural cast of manus and pes prints on a mudstone slab (50 prints).
_____________________________________________________________________________________
Ichnospecies: nipponensis LEE, AZUMA, LEE, SHIBATA & LÜ,
2010
Etymology: In reference to Japan, where the holotype is found.
Holotype: FPDM-F022-1
Locality: Kitadani Locality (Dinosaur Quarry), second footprint horizon, on the Sugiyama River, northern part of the city of Katsuyama, 36°7’N, 136°33’E, Fukui Prefecture, Honshu Island, Japan.
Horizon: Kitadani Formation, Akaiwa subgroup, Tetori Group.
Biostratigraphy:
Age: Barremian Stage, Lower Gallic Subepoch, Lower Early Cretaceous Epoch, Early Cretaceous.
Material:
Material: 5 trackways on a siltstone slab.
_____________________________________________________________________________________
Ichnospecies: dongyangensis CHENG, LÜ, ZHU, AZUMA, ZHENG, JIN,
NODA & SHIBATA,
2013
Etymology:
In reference to Dongyang City of Zhejiang Province, China.
Holotype: DYM-04666-1, 2, 3, 4
Locality: Dongyang, Zhejiang Province, China.
Horizon: Jinhua Formation (once regared as Fangyang Formation).
Biostatigraphy:
Age: Late Cretaceous.
Material: Breakdown as follows;
DYM-04666-1: Manus.
DYM-04666-2: Pes.
DYM-04666-3: Manus.
DYM-04666-4: One block with manus and pes prints.
_____________________________________________________________________________________
Ichnospecies: wuerhoensis LI, WANG & JIANG, 2021
Etymology: In reference to teh locality where the small pterosaur tracks were discovered. The Wuerho region is also teh fossil site of Wuerho Pterosaur fauna.
Holotype: 45MR, 58PR.
Locality: Huangyangquan Reservoir tracksite 1, wuerho District, Karama City, Xinjiang Province, China.
Horizon: Shengjinkou Formation, Tugulo Group.
Biostratigraphy:
Age: Early Cretaceous.
Material: 45MR manus, 58PR pes.
Paratypes:
49ML, 104ML: Manus.
7PR, 29PR: Pes.
Referred material:
1ML-6MR, 8P-28P, 30PR-44ML, 46MR-57ML, 59P-103P, 105ML-114P: 98 tracks.
_____________________________________________________________________________________
Ichnospecies: gracilis HA, KIM, LIM, LOCKLEY, YOO & LIM, 2022
Etymology: Latin, gracilis, "gracile or slender".
Ichnotype: CUE E4P007
Locality: Excavation 4 site, Chungmugong-dong, Jinju Innovation City, Gyeongsangnam-do, South Gyeonsong Province, South Korea.
Horizon: Jinju Formation.
Biostratigraphy:
Age: Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.
Material: Trackway A.
Paratype:
CUE E4007: Trackway B and isolated tacks.