Ichnogenus: Protochirotherium FICHTER & KUNZ, 2004
Etymology: Protochirotherium Proto...in the sense of "Ur".. = Urchirotherium due to the domiant pes digit V.

Species: wolfhagense FICHTER & KUNZ, 2004
Etymology: In reference to Wolfhagen, where the tracks were found.

Holotype: RGMW 10.14.9.1

Locality: Former sandpit in the Stadtwald (Municipal forest) of Wolfhagen near the Stocke Teich (pond), Baden-Wurttenberg State, Germany.

Horizon: Dethfurt Wechselfolge.

Age: Middle Bunter, Early Triassic.

Material: Set of right manus and pes imprents.

Paratypes:

RGMW 10.14.9.7, 10.14.9.4: Tracks.

Referred material:

S 337, S331/VIII, S 330/VII, NMK S 291, NMK S 250.2, NMK S 290.1, NMK S 293, NMK S 250.3, NMK S 296, NMK S 297, NMK S 299, NMK S 298, NMK S 287, NMK S 300.1, NMK S 300.2, NMK S 253, NMK S 251.4, NMK S 251.3, NMK S 251.2, NMK S 7, NMK S 192.1, NMK S 192.2, NMK S 190.1, NMK S 261, NMK S 301, NMK S 276, NMK S 258, NMK S 251.1, NMK S 336, NMK S 244, NMK S 307/6.1, NMK S 207/6.2, NMK S 305/306, NMK S 250.1, SKW 1, SKW 2, SKW 3, SKW 4, SKW 5, SKW 7, SKW 8: Pes and manus tracks.

Note: Some tracks have tail drags and skin impressions.

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Ichnospecies: hauboldi (PTASZYNSKI, 1990) KLEIN, NIEDZWIEDZKI, VOIGT, LAGNANOUI, HMINNA, SABER & SCHNEIDER, 2013
Etymology:
= Chirotherium hauboldi PTASZYNSKI, 1990
= Chiritherium lauboldi PTASZYNSKI, 1990 (sic)
= Brachychirotherium hauboldi (PTASZYNSKI, 1990) PTASZYNSKI, 2000

Syntypes:

Locality: Right bank of Swislina River, Wiory, near Ostrowiec Swietokrzyski, Swietokrzyskie (Holy Cross) Mountains, Poland.

Horizon: Wiory Formation, Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.

Material:

KR 5: 8.1: Deeply impressed, somewhat damaged imprint of right pes.
Note: ZPAL R.7/34: Cast.

KR 9: 14.1: Imprint of right pes with partly preserved digit V.
Note: ZPAL R.7/33: Cast.

KR 1: 56.1: Imprint of right pes.
Note: ZPAL R.7/35: Cast.

MWGUW 01139: 16.1-2: set of right manus-pes imprints somewhat deformed by a sliding movement.

Referred material:

MWGUW 01141: 3.6-5-4: 3 successive pes-manus-pes imprints.

MWGUW 01141: 3.12-28-25-19-24-20-21-23: Irregular trackway.

TP 11.1-2: Set of left pes-manus tracks.

TPW 9. 12.1: Plaster cast of a right pes associated with tail impressions.

TPW 10: 12.24: Plaster cast, imprint of left pes.

TPW 11: 12.80: Plaster cast of right pes.

KR9: 14.2: Partial imprint of left pes probably of the same trackway as the syntype KR 9:14.1.

PTASZYNSKI, 2000

ZPAL R.7/4: 1-11: Well preserved, almost regular track with imprints of all four feet visible although not all completely impressed.

ZPAL R.7/76-83: Track with shallow imprints, made by a large individual.

TP 234: 1, 2, TP 279: 1, 2, TP 463: 1, 2, ZPAL R.7/5: 1, 2, TP 565: 1, 2, TP 1, 2: Isolated sets of pes and manus prints.

TP 12: 24, TP 12: 80, TP 239: 1, TP 468: 1, TP 476: 1, TP 485: 1, TP 574: 1, TP 587B: 1, TP 589: 1: Isolated pes prints.

KLEIN, & NIEDZWIEDZKI, 2012

MPT.P.W1: Trackways no. 1, 4 sets.

MTP.P.W2: Trackways no. 3, 4 sets.

MTP.P.W3: Trackways no. 5-6, 8-10: Partial trackways and isoalted sets with skin texture.

MPT.P.W4: 2 trachways with 4, 5 successive sets respectively, partial trackways and isolated sets with skin texture.

MPT.P.W8: Several isolated sets.

MPT.P.W10: W 26 several isolated concave epipreliefs with skin texture.

MPT.P.W11: Isolated pes of juvenile.

MTP.P.W12, W 14, W 17: Several isoalted large imprints.

MTP.P.W13, W 16, W 18, W 21, W 22, W 27, W 28, W 29 (with skin texture), W 30-W 32, W 33-W36: Isolated sets and partial trackways.

MTP.P.W38, W39: Trackways.

MPT.P.W40: Several isoalted sets

MPT.P.W41-W43: Isolated sets.

MPT.P.W44: Trackway (4 sets), isolated set.

MPT.P.W45, W46 with skin texture: Several isolated sets.

MPT.P.W49-W53: Isolated sets and partial trackway of small individuals.

Unnumbered trackway: 3 succissive sets, partial trackways and isolated sets.

 

= Isochirotherium sanctacrucense PTASYNZKI, 1990
Etymology:

Holotype: MWGUW 01141: 3.29

Locality: Right bank of Swislina River, Wiory, near Ostrowiec Swietokrzyski, Swietokrzyskie (Holy Cross) Mountains, Poland.

Horizon: Wiory Formation, Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.

Material: Complete imprint of right pes, slightly deformed by sliding movement.

Paratype: MWGUW 01140: 13.7: Plaster cast of a well preserved (except of digit IV), of left pes.
Note: ZPAL R. 7/37: Cast.

Referred material:

MWGUW 01141: 3.30-35-17-13-14: With holotype: irregular partly and poorly preserved trackway.

TPW 12: 89.1-2: Set of right manus-pes imprints.

TPW 13.13.17: Plaster cast, partly and poorly impressed imprint of right pes, probably made by the same animal of the paratype.

PTASYNZKI, 2000

ZPAL R.7/17: 1-5: fragmentary track consisting of 2 sets and next consective pes print mostly partially and shallowly impressed, poorly preserved.

ZPAL R.7/15: 1, 3: Isolated sed of right pes and manus.

= Brachychirotherium wiorense PTASZYNSKI, 2000
Etymology: In referene to te type locality.

Holotype: ZPAL R.7/6: 1, 2

Locality: Right bank of Swislina River, Wiory, near Ostrowiec Swietokrzyski, Swietokrzyskie (Holy Cross) Mountains, Poland.

Horizon: Wiory Formation, Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.

Material: Set of right pes and manus.

Referred material

ZPAL R.7/7a: 1: Right manus print.

ZPAL R.7/7b: 2, 3: Left set of pes and manus print.

ZPAL R.7/7c: 1: Right pes print.

TP 50: 1-7: Irregular track composed of right pes imprint and 3 consecutive sets of pes and manus prints.
Note: ZPAL R.7/9: 1-7, cast. Partly destroyed.

TP 5: 1 (not collected): Partialy preserved imprint of left pes.

ZPAL R.7/47: Plaster cast, Isolated left pes digit group, I-IV print.

ZPAL R.7/43: Plaster cast, Isolated print of right pes.

ZPAL R.7/48: 1, 2: Plaster cast, set of right pes and manus.

TP 261: 1,3: Partially preserved set of righ pes and manus.

ZPAL R.7/8: 1, 2: Set of right pes and manus.

ZPAL R.7/13: 6, 7: Poorly preserved set of left pes and manus.

 

= Isochirotherium gierlinskii PTASZYNSNKI, 2000
Etymology: In honor of Gerard Gierlinski, researcher of Mesozoic vertebrate footprints.

Holotype: ZPAL R.7/10: 1, 2

Locality: Right bank of Swislina River, Wiory, near Ostrowiec Swietokrzyski, Swietokrzyskie (Holy Cross) Mountains, Poland.

Horizon: Wiory Formation, Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.

Material: Set of left pes and manus. Forming a natural cast of specimen ZPAL R.7/13: 1, 2

Referred material:

ZPAL R.7/11:1-4: 2 consecutive sets of pes and manus.

ZPAL R.7/14: 1-3: Set of pes and manus and next consecutive pes print.

ZPAL R.7/45:2: Isolated pes print.

ZPAL R.7/13:1-5: Track consisting of 2 sets of pes and manus and the next consecutive pes print.

 

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