Ichnogenus: Brachychirotherium BEURLEN, 1950
Etymology: Greek, brachys, “short” + Chirotherium.

Ichnospecies: gallicum BORNEMANN, 1887 emed. DEMATHIEU, 1974
Etymology: Gallicus, the Roman name for France.

Holotype:

Locality: Eastern France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material:

Referred material:

GAND, HORRA, GALAN-ABELLAN, LOPEZ-GOMEZ, BARRENECHEA, ARCHE, & BENITO, 2010

Locailty: Iberian Ranges, Spain.

Horizon: Landete Formation, Camarena section.

Biostratigaphy:

Age: Base of Muschelkalk, Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

In situ: Manus and pes prints.


_____________________________________________________________________________________

Ichnospecies: parvum (C. H. HITCHCOCK, 1889)
Etymology: Latin, parvus, "small."
= Chirotherium parvum (C. H. HITCHCOCK, 1889)
= Otozoum parvum C. H. HITCHCOCK, 1889

Holotype: Lafayett College, Easton, PA, LC S490

Locality: Quarries at Milford, New Jersey side of the Delaware River, about 30 miles above Trenton, Bergen County, New Jersey .

Horizon: Perkaise Member, Passic Formation.

Biostratigraphy:

Age: Norian Stage, Late Triassic Epoch, Late Triassic.

Material: A large right manus-pes set, the missing posterior half of metatarso-phalangeal pad V is preserved on MCZ 212.

Hypodigm:

LC S488 and MCZ 211: Probably represent 2 individuals of similar size.

 


Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.

Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.

Biostratigraphy:

Age: Late Triassic.

Material:

MCZ 209, 210, AMNH 2257:

 


AVANZINI, 1999


Locality: Upper Val di Non, Val d’Adige, Bolzano, Northern Italy.

Horizon:

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given: Trackways.

 

 

= Brachychirotherium aff. parvium AVANZINI, 1999

Locality: Upper Val di Non, Val d’Adige, Bolzano, Northern Italy.

Horizon:

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.

Material:

Number: Not given: Trackways.

 

 

DEMATHIEU & SAMANA, 1970

Locality: Region of Largentiere, Ardeche Department, France.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material:

Number: Not given: Pes and manus prints.

 

HMINNA, VOIGT, KLEIN, SABER, SCHNEIDER & HMICH, 2013

Locality: CDUE locality 31 (N33°08'57.3", W008°08'42.2"), Sidi Said Maachou area, Western Meseta, Morocco.

Horizon: Machrra Abbas Member, Upper part of the Oued Oum Er Rbiaa Formation.

Biostratigraphy:

Age: Late Triassic.

Material:

CDUE 5 (part and counterpart): Right pes and manus.

CDUE 6 (part and counterpart): Right pes and manus.

CDUE 7: Left pes.

Locality: CDUE locality 42 (N33°08'53.9", W008°07'54.2"), Sidi Said Maachou area, Western Meseta, Morocco.

Horizon: Machrra Abbas Member, Upper part of the Oued Oum Er Rbiaa Formation.

Biostratigraphy:

Age: Late Triassic.

Material:

CDUE 351: Poorly preserved fragmentary left pes.

 

LUCAS, SPIELMANN, KLEIN & LERNER, 2010

Locality: NMMNH L-472, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: Quay Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-22418, P-22419: Manus and pes prints.

 

Locality: NMMNH L-1477, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: Duke Ranch Member, Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-22424: Manus and pes prints.

NMMNH P-22425: Concave epirelief.

 

Locality: NMMNH L-5775, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-44177, P-44178, P-44179, P-44184: Manus and pes prints.

NMMNH P-44179: Concave epirelief.

 

Locality: NMMNH L-5791, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH-P 44190: Manus and pes prints.

 

Locality: NMMNH L-5792, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH-P-44191-P-44199: Manus and pes prints.

 

Locality: NMMNH L-5793, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-44195-P-44199, P-44201, P-44203: Manus and pes prints.

 

Locality: NMMNH L-5794, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-44204-P-44212: Manus and pes prints.

 

Locality: NMMNH L-5795, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-44213-P-44215: Manus and pes prints.

 

Locality: NMMNH L-5798, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-44217, P-44219, P-44222: Manus and pes prints.

 

Locality: NMMNH L5799, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-44224-P-44226: Manus and pes prints.

 

Locality: NMMNH L-7287, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-57443, 57445: Concave epirelief.

Locality: NMMNH L-7288, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-57441, P-59000: Manus and pes prints.

 

Locality: NMMNH L-7290, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-57434: Manus and pes prints.

 

Locality: NMMNH L-7574, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-57477: Manus and pes prints.

 

Locality: NMMNH L-7768, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.

Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.

Biostratigraphy:

Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.

Material:

NMMNH P-58998, P-58999: Manus and pes prints.

 

= Brachychirotherium cf. B. parvum SUES, & OLSEN, 2015

Locality: Locality B and C, Cliffs and beach just south-southwest of Paddy Island, Kings County, Nova Scotia Province, Canada.

Horizon: Red Head Member, lower Blomidon Formation, Newark Supergroup.

Biostratigraphy:

Age: Early Norian Stage, Lower-Middle Late Triassic Epoch, Late Triassic.

Material:

In situ: Manus and pes prints.

 

WEEMS, 2022

Locality: Main track surface in Valley Quarries Fairfield Operation, near Fairfield,Gettysburg Basin, Adams County, Pennsylvania.

Horizon: Passaic Formation, Newark Super Group.

Biostratigraphy:

Age: Upper Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

CM 92829, 92832, 92834, insitu: Trackways.

WEEMS, & KIMMEL, 1993

Locality: Manassas 7.5'-quadrangle along Compton Road, east of Virgina Route 28, Culpeper Basin, Fairfax County, Virgina.

Horizon: Lower Poolsveille Member, Manassas Sandstone.

Biostratigraphy:

Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material:

USNM 412534 (part and counterpart): Left manus and pes.

 


= Chirotherium copei BOCK, 1952
Etymology: In honor of Edward Drinker Cope.

Holotype: ANS 15210

Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.

Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.

Biostratigraphy:

Age: Late Triassic.

Material:

_____________________________________________________________________________________

Ichnospecies: hessei (SOERGEL, 1925)
Etymology:
= Chirotherium hessei SOERGEL, 1925

Holotype:

Locality: Kauerndorf near Kulmbach, Bayern (Baveria) State, Germany.

Horizon:

Biostratigraphy:

Age:?

Material: Obscure manus and pes prints.
_____________________________________________________________________________________

Ichnospecies: ischigualastianus (HUENE, 1931) HUNT & LUCAS, 2007
= Rigalites ischigualastianus HUENE, 1931
Etymology: In reference to Ischigualasto, San Juan Province, Argentina.

Neotype: PVL 2610

Locality: Quebrada de los Rastros, Agua de la Pena, 29°30'S, 69°00'W, Ischigualasto, San Juan Province, Argentina.

Horizon: Los Rastros Formation.

Biostratigraphy:

Age: Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.

Material: Footprints.

_____________________________________________________________________________________
Ichnospecies: thuringiacum (RUHLE von LILIENSTERN, 1939)
Etymology: in reference to Thuringia State, Germany.
= Chirotherium thuringiacum RUHLE von LILIENSTERN, 1938

Lectotype: MB 1969.54.244

Locality: 1 km SSW of Dorfes Rudelsdorf, (Blatt Rodach), South Thuringia State, Germany.

Horizon: Blasensandstein, Middle Keuper.

Biostratigraphy:

Age: Late Triassic.

Material: Footprints of manus and pes.

Referred material:

KARL & HAUBOLD, 1998

Locality: North Bayern (Bavaria), Germany.

Horizon: Coburger Sandstein, Middle Keuper.

Biostratigraphy:

Age: Carnian/Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

UEN F3, 54:50, F5, F6:49, F7:51, F8:34, F10, F12:44, F14:8, F15:55, F19:5, F20:?20, F21, F23, F25, F59, F60:2, F61”40, F62”43, F63, P64”28, F65, F79, F80, F81, F82, MNKB F87:62, MNKB o. Nr, NMC 17912a, 17912b, NMC 8758, NMC L. 124, NHMS VT 99, NHMS VT100: Tracks and trackways.

 

PETTI, AVANZINI, NICOSIA, GIRARDI, BERNARDI, FERRETTI, SCHIROLLI & DAL SASSO, 2009

Locality: Zone Village, about 4 km east of the north-eastern side fo the Lake Iseo, Southern Alps, Brescia, Lombardy, Northenr Italy.

Horizon: Val Sabbia Sandstone.

Biostratigraphy:

Age: Middle Carnian Stage, Late Triassic Epoch, Early Late Triassic.

Material:

2 levels of trackways.

Level 1

BsZ-A: 10 manus-pes prints.

BsZ-B: About 9 manus-pes prints.

BsZ-C: Poorly preserved footprints.

 

Level 2

BsZ-D: 4 well preserved manus-pes prints.

BsZ-E: Manus-pes prints.

BsE-F: Manus-pes prints.


WEEMS, 2022

Locality: Main track surface in Valley Quarries Fairfield Operation, near Fairfield,Gettysburg Basin, Adams County, Pennsylvania.

Horizon: Passaic Formation, Newark Super Group.

Biostratigraphy:

Age: Upper Norian Stage, Late Triassic Epoch, Late Triassic.

Material:

CM 92835, 92836, in situ: Trackways.

= Brachychirotherium coburgense AUMANN, 1957
Etymology:

Holotype: NMC o Nr

Locality: Near Coburg, Germany.

Horizon: Lower Burgsandstein.

Biostratigraphy:

Age:

Material:

Referred material:

KARL & HAUBOLD, 1998

Kategorie 1

MB 1969.54.244, NMC o Nr., UEN F1:38b, F3, F13:38a, F14: 18, F65, MNKB F87:62, MNKB o. Nr., NMC 17912a, NMC 17912b, NHMS VT 100: Tracks.

Kategorie 2

UEN F2, F4:50, F5, F6:49, F7:51, F8:34, F10, F11, F12:44, F23, F25, F27, F60:2, F61:40, F62:43, F63, F64:28, F79, F82, NMC 8758, NMC L 124, NHMS VT 99, MB o. Nr: Tracks.

Kategore 3

MB 1969.54.232, MB 1969.54.233, MB 1969.54, 234, MB 1969.54.235, MB 1969.54.239, MB 1969.54.240, MB 1969.54.241, MB 1969.54.243, UEN F15:55, F16, F17, F18, F19:5, F20:?20, UEN F21, UEN F26, UEN F28, F20, F30, F59, F80, F81, F83: Tracks.


= Pseudotetrasauropus dulcis ELLENBERGER, 1972
Etymology:
= Brachychirotherium dulcis (ELLENBERGER, 1972) OLSEN & GALTON, 1984

Holotype:

Locality: Sandstone of Seaka, South Africa.

Horizon: Lower Stormberg Assemblage.

Biostratigraphy:

Age: Late Triassic.

Material: Manus and pes prints.

_____________________________________________________________________________________
Ichnospecies: hassfurtense BEURLEN, 1950
Etymology: In reference to Haßurt, North Bayern (Bavaria),Germany.

Lectotype: UEN F22:1

Locality: Haßurt (Main), and Zeil-Ebelsbach, North Bayern (Bavaria),Germany.

Horizon: Middle Keuper.

Biostratigraphy:

Age: Late Triassic.

Material: Pes and manus tracks.

Paratypes:

UEN F9, UEN F24: Pes and manus tracks.

_____________________________________________________________________________________
Ichnospecies: eyermani (BAIRD, 1957) HAUBOLD, 1971
Etymology:
= Chirotherium eyermani BAIRD, 1957

Holotype: MCZ 134

Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.

Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.

Biostratigraphy:

Age: Late Triassic.

Material: An isolated left pes imprint.

Referred material:

OLSEN & FLYNN, 1989

Locality: Nyack Beach State Park, Haverstraw, Rockland County, New York.

Horizon: Stockton Formation.

Biostratigraphy:

Age: Late Triassic.

Material:

YPM 7731: Pes overlapping a manus-pes of Apatopus lineatus.


_____________________________________________________________________________________

Ichnospecies: circaparvum DEMATHIEU, 1967
Etymology:

Holotype:

Locality: Chassely, Mont d’Or lyonnas, Rhone Department, France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material: 5 manus and 6 pes.

Referred material:

Localities: Montagne de Crussol, Pont d’Argent, Ardeche Department, France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material:

Number: Not given:

 

= Brachychirotherium cf. circaparvum DEMATHIEU & OOSTERNIK, 1983

Locality: Winterswijk, eastern part of Holland.

Horizon: Lower Muschelkalk.

Biostratigraphy:

Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.

Material:

Platte D151: Manus and pes print.

_____________________________________________________________________________________

Ichnospecies: tintanti DEMATHIEU, 1967
Etymology: In honor of Professor H. Tintant.

Holotype:

Locality: Chasselay, Mont d’Or lyonnais, Rhone Department, France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material: 6 prints.
_____________________________________________________________________________________

Ichnospecies: harrasense (HAUBOLD, 1967)
Etymology:
= Chirotherium harrasense HAUBOLD, 1967

Holotype:

Locality: Near Hildburghausen, Thuringia State, Germany.

Horizon:

Biostratigraphy:

Age: Late Buntsandstein, Scythian Epoch, Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: paraeparvum (HAUBOLD, 1967)
Etymology:
= Chirotheirum paraeparvum HAUBOLD, 1967

Holotype:

Locality: Near Hildburghausen, Thuringia State, Germany.

Horizon:

Biostratigraphy:

Age: Late Buntsandstein, Scythian Epoch, Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: lorteti (HAUBOLD, 1970)
Etymology:
= Chirotherium lorteti HAUBOLD, 1970

Holotype:

Locality: Vincelles, Senecey-le-Grand, Saone-et-Loire Department, France.

Horizon:

Biostratigraphy:

Age: Middle Triassic.

Material:
_____________________________________________________________________________________


Ichnospecies: minor (ELLENBERGER, 1970)
Etymology: Latin, minor, "minor, small."
= Deuterosauropodopus minor ELLENBERGER, 1970

Holotype:

Locality: Subeng, Lesotho, Southern Africa.

Horizon: Lower Stormberg Assemblage.

Biostratigraphy:

Age: Late Triassic.

Material: Manus and pes prints.

Plesiotypes: 3 prints.
_____________________________________________________________________________________

Ichnospecies: seakaensis (ELLENBERGER, 1970)
Etymology:
= Tetrasauropos seakaensisELLENBERGER, 1970

Holotype:

Locality: Lesotho, Southern Africa.

Horizon: Lower Stormberg Assemblage.

Biostratigraphy:

Age: Late Triassic.

Material:
_____________________________________________________________________________________

Ichnospecies: pachydactylum DEMATHIEU & GAND, 1975
Etymology:

Holotype:

Locality: Saint-Sernin-du-Bios, Saone-et-Loire Department, France.

Horizon:

Biostratigraphy:

Age: Late Triassic.

Material: Trackways.

_____________________________________________________________________________________

Ichnospecies: sedunensis (DEMATHIEU & WEIDMANN, 1982)
Etymology:
= Deuterodauropodopus sedunensis DEMATHIEU & WEIDMANN, 1982

Holotype: Empreinte no 26

Locality: Aiguilles Rouges, Vieux Emosson, Finhaut, Valais, France.

Horizon:

Biostratigraphy:

Age: Late Ladinian Stage, Upper Mid Triassic Epoch or Carnian Stage, Lower Late Triassic Epoch, Late Triassic.

Material: Tetradactylus prints.

Hypodigm:

Empreintes nos 8, 12, 13, 26, 26B, 35, 38, 40, 42, 43, 55, 105:

_____________________________________________________________________________________

Ichnospecies: kuhni DEMATHIEU & HAUBOLD, 1982
Etymology:

Holotype:

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

Referred material:

PTASYNSKI, 1990 in FUGLEWICZ, PTASYNSKI & RDZANEK, 1990

Locality: Wiory near Ostrowiec Swietokrzyski, Swietokrzyskie Mountains, Poland.

Horizon: Labyrinthodontid beds.

Biostratigraphy:

Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.

Material:

5.1: Partly preserved imprint of left pes.

TPW 15: 13.14: Plaster cast of left pes digits group I-IV.

TPW 16: 39.1: Imprint of right pes.

TPW 17: 57.1, TPW 18: 57.5: Plaster casts, set of right manus-pes imprints.

KR 12: 44.1-2: Poor partly preserved set of manus-pes imprints.

_____________________________________________________________________________________
Ichnospecies: paneneparvum DEMATHIEU & LEITZ, 1982
Etymology:

Holotype:

Locality: Franconia, Germany.

Horizon:

Biostratigraphy:

Age: Early Triassic.

Material:

_____________________________________________________________________________________