Ichnogenus: Brachychirotherium BEURLEN, 1950
Etymology:
Greek, brachys, “short” + Chirotherium.
Ichnospecies: gallicum BORNEMANN, 1887 emed. DEMATHIEU, 1974
Etymology: Gallicus, the Roman name for France.
Holotype:
Locality: Eastern France.
Horizon:
Biostratigraphy:
Age: Middle Triassic.
Material:
Referred material:
GAND, HORRA, GALAN-ABELLAN, LOPEZ-GOMEZ, BARRENECHEA, ARCHE, & BENITO, 2010
Locailty: Iberian Ranges, Spain.
Horizon: Landete Formation, Camarena section.
Biostratigaphy:
Age: Base of Muschelkalk, Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material:
In situ: Manus and pes prints.
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Ichnospecies: parvum (C. H. HITCHCOCK, 1889)
Etymology: Latin, parvus, "small."
= Chirotherium parvum (C. H. HITCHCOCK, 1889)
= Otozoum parvum C. H. HITCHCOCK, 1889
Holotype: Lafayett College, Easton, PA, LC S490
Locality: Quarries at Milford, New Jersey side of the Delaware River, about 30 miles above Trenton, Bergen County, New Jersey .
Horizon: Perkaise Member, Passic Formation.
Biostratigraphy:
Age: Norian Stage, Late Triassic Epoch, Late Triassic.
Material: A large right manus-pes set, the missing posterior half of metatarso-phalangeal pad V is preserved on MCZ 212.
Hypodigm:
LC S488 and MCZ 211: Probably represent 2 individuals of similar size.
Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.
Biostratigraphy:
Age: Late Triassic.
Material:
MCZ 209, 210, AMNH 2257:
AVANZINI, 1999
Locality: Upper Val di Non, Val d’Adige, Bolzano, Northern Italy.Horizon:
Biostratigraphy:
Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material:
Number: Not given: Trackways.
= Brachychirotherium aff. parvium AVANZINI, 1999
Locality: Upper Val di Non, Val d’Adige, Bolzano, Northern Italy.
Horizon:
Biostratigraphy:
Age: Anisian Stage, Lower Mid Triassic Epoch, Middle Triassic.
Material:
Number: Not given: Trackways.
DEMATHIEU & SAMANA, 1970
Locality: Region of Largentiere, Ardeche Department, France.
Horizon:
Biostratigraphy:
Age: Late Triassic.
Material:
Number: Not given: Pes and manus prints.
HMINNA, VOIGT, KLEIN, SABER, SCHNEIDER & HMICH, 2013
Locality: CDUE locality 31 (N33°08'57.3", W008°08'42.2"), Sidi Said Maachou area, Western Meseta, Morocco.
Horizon: Machrra Abbas Member, Upper part of the Oued Oum Er Rbiaa Formation.
Biostratigraphy:
Age: Late Triassic.
Material:
CDUE 5 (part and counterpart): Right pes and manus.
CDUE 6 (part and counterpart): Right pes and manus.
CDUE 7: Left pes.
Locality: CDUE locality 42 (N33°08'53.9", W008°07'54.2"), Sidi Said Maachou area, Western Meseta, Morocco.
Horizon: Machrra Abbas Member, Upper part of the Oued Oum Er Rbiaa Formation.
Biostratigraphy:
Age: Late Triassic.
Material:
CDUE 351: Poorly preserved fragmentary left pes.
LUCAS, SPIELMANN, KLEIN & LERNER, 2010
Locality: NMMNH L-472, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: Quay Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-22418, P-22419: Manus and pes prints.
Locality: NMMNH L-1477, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: Duke Ranch Member, Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-22424: Manus and pes prints.
NMMNH P-22425: Concave epirelief.
Locality: NMMNH L-5775, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-44177, P-44178, P-44179, P-44184: Manus and pes prints.
NMMNH P-44179: Concave epirelief.
Locality: NMMNH L-5791, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH-P 44190: Manus and pes prints.
Locality: NMMNH L-5792, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH-P-44191-P-44199: Manus and pes prints.
Locality: NMMNH L-5793, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-44195-P-44199, P-44201, P-44203: Manus and pes prints.
Locality: NMMNH L-5794, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-44204-P-44212: Manus and pes prints.
Locality: NMMNH L-5795, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-44213-P-44215: Manus and pes prints.
Locality: NMMNH L-5798, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-44217, P-44219, P-44222: Manus and pes prints.
Locality: NMMNH L5799, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-44224-P-44226: Manus and pes prints.
Locality: NMMNH L-7287, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-57443, 57445: Concave epirelief.
Locality: NMMNH L-7288, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-57441, P-59000: Manus and pes prints.
Locality: NMMNH L-7290, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-57434: Manus and pes prints.
Locality: NMMNH L-7574, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-57477: Manus and pes prints.
Locality: NMMNH L-7768, northeast edge of Llano Estacado, Guadalupe and Quay Counties, east-central New Mexico.
Horizon: San Jon Creek Member, Redonda Formation, Chinle Group.
Biostratigraphy:
Age: Apachean, Late Norian Stage- Rhaetian Stage, Upper Late Triassic Epoch, Late Late Triassic.
Material:
NMMNH P-58998, P-58999: Manus and pes prints.
= Brachychirotherium cf. B. parvum SUES, & OLSEN, 2015
Locality: Locality B and C, Cliffs and beach just south-southwest of Paddy Island, Kings County, Nova Scotia Province, Canada.
Horizon: Red Head Member, lower Blomidon Formation, Newark Supergroup.
Biostratigraphy:
Age: Early Norian Stage, Lower-Middle Late Triassic Epoch, Late Triassic.
Material:
In situ: Manus and pes prints.
WEEMS, 2022
Locality: Main track surface in Valley Quarries Fairfield Operation, near Fairfield,Gettysburg Basin, Adams County, Pennsylvania.
Horizon: Passaic Formation, Newark Super Group.
Biostratigraphy:
Age: Upper Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
CM 92829, 92832, 92834, insitu: Trackways.
WEEMS, & KIMMEL, 1993
Locality: Manassas 7.5'-quadrangle along Compton Road, east of Virgina Route 28, Culpeper Basin, Fairfax County, Virgina.
Horizon: Lower Poolsveille Member, Manassas Sandstone.
Biostratigraphy:
Age: Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material:
USNM 412534 (part and counterpart): Left manus and pes.
= Chirotherium copei BOCK, 1952
Etymology: In honor of Edward Drinker Cope.Holotype: ANS 15210
Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.
Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.
Biostratigraphy:
Age: Late Triassic.
Material:
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Ichnospecies: hessei (SOERGEL, 1925)
Etymology:
= Chirotherium hessei SOERGEL, 1925
Holotype:
Locality: Kauerndorf near Kulmbach, Bayern (Baveria) State, Germany.
Horizon:
Biostratigraphy:
Age:?
Material: Obscure manus and pes prints.
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Ichnospecies: ischigualastianus (HUENE, 1931) HUNT & LUCAS, 2007
= Rigalites ischigualastianus HUENE, 1931
Etymology: In reference to Ischigualasto, San Juan Province, Argentina.
Neotype: PVL 2610
Locality: Quebrada de los Rastros, Agua de la Pena, 29°30'S, 69°00'W, Ischigualasto, San Juan Province, Argentina.
Horizon: Los Rastros Formation.
Biostratigraphy:
Age: Ladinian Stage, Upper Mid Triassic Epoch, Middle Triassic.
Material: Footprints.
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Ichnospecies: thuringiacum (RUHLE
von LILIENSTERN, 1939)
Etymology: in reference to Thuringia State,
Germany.
= Chirotherium thuringiacum RUHLE von
LILIENSTERN, 1938
Lectotype: MB 1969.54.244
Locality: 1 km SSW of Dorfes Rudelsdorf, (Blatt Rodach), South Thuringia State, Germany.
Horizon: Blasensandstein, Middle Keuper.
Biostratigraphy:
Age: Late Triassic.
Material: Footprints of manus and pes.
Referred material:
KARL & HAUBOLD, 1998
Locality: North Bayern (Bavaria), Germany.
Horizon: Coburger Sandstein, Middle Keuper.
Biostratigraphy:
Age: Carnian/Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
UEN F3, 54:50, F5, F6:49, F7:51, F8:34, F10, F12:44, F14:8, F15:55, F19:5, F20:?20, F21, F23, F25, F59, F60:2, F61”40, F62”43, F63, P64”28, F65, F79, F80, F81, F82, MNKB F87:62, MNKB o. Nr, NMC 17912a, 17912b, NMC 8758, NMC L. 124, NHMS VT 99, NHMS VT100: Tracks and trackways.
PETTI, AVANZINI, NICOSIA, GIRARDI, BERNARDI, FERRETTI, SCHIROLLI & DAL SASSO, 2009
Locality: Zone Village, about 4 km east of the north-eastern side fo the Lake Iseo, Southern Alps, Brescia, Lombardy, Northenr Italy.
Horizon: Val Sabbia Sandstone.
Biostratigraphy:
Age: Middle Carnian Stage, Late Triassic Epoch, Early Late Triassic.
Material:
2 levels of trackways.
Level 1
BsZ-A: 10 manus-pes prints.
BsZ-B: About 9 manus-pes prints.
BsZ-C: Poorly preserved footprints.
Level 2
BsZ-D: 4 well preserved manus-pes prints.
BsZ-E: Manus-pes prints.
BsE-F: Manus-pes prints.
WEEMS, 2022
Locality: Main track surface in Valley Quarries Fairfield Operation, near Fairfield,Gettysburg Basin, Adams County, Pennsylvania.
Horizon: Passaic Formation, Newark Super Group.
Biostratigraphy:
Age: Upper Norian Stage, Late Triassic Epoch, Late Triassic.
Material:
CM 92835, 92836, in situ: Trackways.
= Brachychirotherium coburgense AUMANN, 1957
Etymology:Holotype: NMC o Nr
Locality: Near Coburg, Germany.
Horizon: Lower Burgsandstein.
Biostratigraphy:
Age:
Material:
Referred material:
KARL & HAUBOLD, 1998
Kategorie 1
MB 1969.54.244, NMC o Nr., UEN F1:38b, F3, F13:38a, F14: 18, F65, MNKB F87:62, MNKB o. Nr., NMC 17912a, NMC 17912b, NHMS VT 100: Tracks.
Kategorie 2
UEN F2, F4:50, F5, F6:49, F7:51, F8:34, F10, F11, F12:44, F23, F25, F27, F60:2, F61:40, F62:43, F63, F64:28, F79, F82, NMC 8758, NMC L 124, NHMS VT 99, MB o. Nr: Tracks.
Kategore 3
MB 1969.54.232, MB 1969.54.233, MB 1969.54, 234, MB 1969.54.235, MB 1969.54.239, MB 1969.54.240, MB 1969.54.241, MB 1969.54.243, UEN F15:55, F16, F17, F18, F19:5, F20:?20, UEN F21, UEN F26, UEN F28, F20, F30, F59, F80, F81, F83: Tracks.
= Pseudotetrasauropus dulcis ELLENBERGER, 1972
Etymology:
= Brachychirotherium dulcis (ELLENBERGER, 1972) OLSEN & GALTON, 1984Holotype:
Locality: Sandstone of Seaka, South Africa.
Horizon: Lower Stormberg Assemblage.
Biostratigraphy:
Age: Late Triassic.
Material: Manus and pes prints.
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Ichnospecies: hassfurtense BEURLEN, 1950
Etymology: In reference to Haßurt, North Bayern (Bavaria),Germany.
Lectotype: UEN F22:1
Locality: Haßurt (Main), and Zeil-Ebelsbach, North Bayern (Bavaria),Germany.
Horizon: Middle Keuper.
Biostratigraphy:
Age: Late Triassic.
Material: Pes and manus tracks.
Paratypes:
UEN F9, UEN F24: Pes and manus tracks.
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Ichnospecies: eyermani (BAIRD, 1957) HAUBOLD, 1971
Etymology:
= Chirotherium eyermani BAIRD, 1957
Holotype: MCZ 134
Locality: Smith Clark quarry near Milford, 1.25 km north of Delaware River Bridge, Hunterdon County, New Jersey.
Horizon: Lower Passic Formation (Old Upper Brunswick Formation), about 5100 meters above the Triassic Base, Level B, Newark Series.
Biostratigraphy:
Age: Late Triassic.
Material: An isolated left pes imprint.
Referred material:
OLSEN & FLYNN, 1989
Locality: Nyack Beach State Park, Haverstraw, Rockland County, New York.
Horizon: Stockton Formation.
Biostratigraphy:
Age: Late Triassic.
Material:
YPM 7731: Pes overlapping a manus-pes of Apatopus lineatus.
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Ichnospecies: circaparvum DEMATHIEU, 1967
Etymology:
Holotype:
Locality: Chassely, Mont d’Or lyonnas, Rhone Department, France.
Horizon:
Biostratigraphy:
Age: Middle Triassic.
Material: 5 manus and 6 pes.
Referred material:
Localities: Montagne de Crussol, Pont d’Argent, Ardeche Department, France.
Horizon:
Biostratigraphy:
Age: Middle Triassic.
Material:
Number: Not given:
= Brachychirotherium cf. circaparvum DEMATHIEU & OOSTERNIK, 1983
Locality: Winterswijk, eastern part of Holland.
Horizon: Lower Muschelkalk.
Biostratigraphy:
Age: Anisian Stage, Lower Mid Triassic Epoch, Early Middle Triassic.
Material:
Platte D151: Manus and pes print.
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Ichnospecies: tintanti DEMATHIEU, 1967
Etymology: In honor of Professor H. Tintant.
Holotype:
Locality: Chasselay, Mont d’Or lyonnais, Rhone Department, France.
Horizon:
Biostratigraphy:
Age: Middle Triassic.
Material: 6 prints.
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Ichnospecies: harrasense (HAUBOLD, 1967)
Etymology:
= Chirotherium harrasense HAUBOLD, 1967
Holotype:
Locality: Near Hildburghausen, Thuringia State, Germany.
Horizon:
Biostratigraphy:
Age: Late Buntsandstein, Scythian Epoch, Late Triassic.
Material:
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Ichnospecies: paraeparvum (HAUBOLD, 1967)
Etymology:
= Chirotheirum paraeparvum HAUBOLD, 1967
Holotype:
Locality: Near Hildburghausen, Thuringia State, Germany.
Horizon:
Biostratigraphy:
Age: Late Buntsandstein, Scythian Epoch, Late Triassic.
Material:
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Ichnospecies: lorteti (HAUBOLD, 1970)
Etymology:
= Chirotherium lorteti HAUBOLD, 1970
Holotype:
Locality: Vincelles, Senecey-le-Grand, Saone-et-Loire Department, France.
Horizon:
Biostratigraphy:
Age: Middle Triassic.
Material:
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Ichnospecies: minor (ELLENBERGER, 1970)
Etymology: Latin, minor, "minor, small."
= Deuterosauropodopus minor ELLENBERGER, 1970
Holotype:
Locality: Subeng, Lesotho, Southern Africa.
Horizon: Lower Stormberg Assemblage.
Biostratigraphy:
Age: Late Triassic.
Material: Manus and pes prints.
Plesiotypes: 3 prints.
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Ichnospecies: seakaensis (ELLENBERGER, 1970)
Etymology:
= Tetrasauropos seakaensisELLENBERGER, 1970
Holotype:
Locality: Lesotho, Southern Africa.
Horizon: Lower Stormberg Assemblage.
Biostratigraphy:
Age: Late Triassic.
Material:
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Ichnospecies: pachydactylum DEMATHIEU & GAND, 1975
Etymology:
Holotype:
Locality: Saint-Sernin-du-Bios, Saone-et-Loire Department, France.
Horizon:
Biostratigraphy:
Age: Late Triassic.
Material: Trackways.
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Ichnospecies: sedunensis (DEMATHIEU & WEIDMANN,
1982)
Etymology:
= Deuterodauropodopus sedunensis DEMATHIEU & WEIDMANN,
1982
Holotype: Empreinte no 26
Locality: Aiguilles Rouges, Vieux Emosson, Finhaut, Valais, France.
Horizon:
Biostratigraphy:
Age: Late Ladinian Stage, Upper Mid Triassic Epoch or Carnian Stage, Lower Late Triassic Epoch, Late Triassic.
Material: Tetradactylus prints.
Hypodigm:
Empreintes nos 8, 12, 13, 26, 26B, 35, 38, 40, 42, 43, 55, 105:
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Ichnospecies: kuhni DEMATHIEU & HAUBOLD, 1982
Etymology:
Holotype:
Locality:
Horizon:
Biostratigraphy:
Age:
Material:
Referred material:
PTASYNSKI, 1990 in FUGLEWICZ, PTASYNSKI & RDZANEK, 1990
Locality: Wiory near Ostrowiec Swietokrzyski, Swietokrzyskie Mountains, Poland.
Horizon: Labyrinthodontid beds.
Biostratigraphy:
Age: Middle Buntsandstein, Scythian Epoch, Early Triassic.
Material:
5.1: Partly preserved imprint of left pes.
TPW 15: 13.14: Plaster cast of left pes digits group I-IV.
TPW 16: 39.1: Imprint of right pes.
TPW 17: 57.1, TPW 18: 57.5: Plaster casts, set of right manus-pes imprints.
KR 12: 44.1-2: Poor partly preserved set of manus-pes imprints.
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Ichnospecies:
paneneparvum DEMATHIEU & LEITZ, 1982
Etymology:
Holotype:
Locality: Franconia, Germany.
Horizon:
Biostratigraphy:
Age: Early Triassic.
Material:
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