Genus: Spinosaurus STROMER, 1915
Etymology: Latin, spina, “spine” and Greek, sauros, “lizard”: "Spine lizard."

Species: aegyptiacus STROMER, 1915
Etymology: In reference to the country it was found, Egypt.

Holotype: BSP 1912 VII 19 (IPHG 1912 VIII 19) (Destroyed during WWII).

Locality: 3 km north of the Gebel el Dist in the basin of Baharija Oasis, Egypt.

Horizon: Baharija Formation.

Biostratigraphy:

Age: ?Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowemrost Gulf Epoch, Middle Cretaceous.

Material: Dentary with teeth, cervical, dorsal, sacral and caudal vertebrae, ribs.

Two verisons of a short skull.

Two versions of a longer skull. The bottom is after a new undescribed skull.

Three possible skeletal reconstructions.

Possible bending of the back (George Olshevsky).

IBRAHIM, SERENO, DAL SASSO, MAGANUCO, FABBRI, MARTILL, ZOUHRI, MYHRVOLD, & IURINO, 2014

Neotype: FSAC-KK 11888

Locality: Spinosaurus site, near Zrigat (also spelled Zrigat), South-east facing escarpemnt fringing the Aferdou Zrigat (Ikhf n Taqmout) plateau, an isoalted mesa of the more extensive Jbel Al Marka Plateau, northern margin of the Tafilalt Basin, Errachidia Province, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: Neural spine, left ilium, right manual II-1 phalanx, left femur, right tibia, and right pedal ungual.

Note: New material as per IBRAHIM, MAGANUCO, DAL SASSO, FABBRI, AUDITORE, BINDELLINI, MARTILL, ZOUHIR, MATTARELLI, UNWIN, WIEMANN, BONADONNA,AMANE, JAKUBCZAK, JOGER, LAUDER 7 PIERCE, 2020: Fragments of cranium and mandibles, several previously missing bones of the left ad right pes, and mostimportantly, most of the caudal vertebrae.

 

Referred material:

BENYOUCEF, CAVIN, MEBARKI, ADACI & BENSALAH, 2015

Locality: Kenasada and Nenaguir, Guir Basin, Algeria.

Horizon: Kem Kem beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

UTL25-MN1-12: 79 teeth.

UTL25-KD1-1: 2 teeth.


GOEDERT, AMIOT, BOUDAD, BUFFETAUT, FOUREL, GODEFROIT, KUSUHASHI, SUTEETHORN, TONG, WATABE, & lECUYER, 2016

Locality: Khetitila Shrira, Morocco.

Horizon: Ifezounae Formation.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Mca: Tooth.

 

HENDRICKX, MATEUS, & BUFFETAUT, 2016

Locality: Kem Kem area, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: 

Morphotype I

MHNM.KK375, KK377, MSNM V6896: 6 quadrates.

 

Locality: Jorf (Tafilalt), northwest of Erfound, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: 

Morphhotype I

MHNM.KK374: Quadrate (found in reddish iron-rich sandstones).

MHNM KK 378: 2 quadrates (found in ironless layers of white to yellow sandstones).

 

= cf. Spinosaurus aegyptiacus LAKIN & LONGRICH, 2019

Locality: Near Begaa, Er Rachidia, Morocco.

Horizon: Kem Kem Beds, Ifezouane Formation.

Biostratigraphy:

Age: Cenomanian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

FASC-KK-18118: Anterior dorsal vertebra.

FSAC-KK-7280: A cervical vertebra.

 

MAGANUCO & DAL SASSO, 2018

Locality: A few kilometers south of Erfound, between the villages of Taouz and Begaa, Errachidia Province, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: 

MSNM V6894: Pedal ungual phalanx of a juvenile.

 

MILNER, 1996, 2001

Locality: Taouz, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material: 

BMNH R 16420: Premaxilla, and fragmentary maxilla.

BMNH R 16421: Left dentary.

 

= Spinosaurus cf. aegyptiacus BUFFETAUT, 1989

Locality: Near the town of Taouz, at the base of a vast plateau known as the Hammada du Guir, Morocco.

Horizon: Continental Red Beds.

Biostratigraphy:

Age: Upper Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch or Lower Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

IMGP 969-3: Tooth.

IMGP 969-1: Fragmentary right maxilla.

 

= Spinosaurus cf. aegyptiacus BUFFETAUT & OUAJA, 2002

Locality: Jebel Miteur, Tataouine Governorate, Southern Tunisia.

Horizon: Chenini Sandstones.

Biostratigraphy:

Age: Late Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

ONM BM 231: Rostral part of left dentary containing 4 alveoli.

 

= Spinosaurus cf. aegyptiacus DAL SASSO, MAGANUCO, BUFFETAUT & MENDEZ, 2005 (published in 2006)

Locality: Kem Kem, east of the town of Taouz, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

MSNM V4047: Snout.

Note: Has an isolated vertebra of ?Onchopristis sp embedded between the right second premaxilary alveolus and its erupting tooth.

 

Locality: Kem Kem beds, N30°02', W5°12', near the outpost in Keneg ed Dal, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

UCPC-2: Caudal part of a pair of fused nasals and a small fragment of a left maxilla.

AZUMA, 2005

FPDM-V7011: Cast of a new undescribed skull.

Number: Not given: Dorsal vertebra.

IBRAHIM, SERENO, DAL SASSO, MAGANUCO, FABBRI, MARTILL, ZOUHRI, MYHRVOLD, & IURINO, 2014

UCRC PV601: Anterior dorsal vertebra.

FSAC-KK 11888: Neural spine, left ilium, right manual II-1 phalanx, left femur, right tibia, and right pedal ungual..

CA30: Mid cuadal vertebrta.

UCRC VP5: Anterior caudal vertebra.

 

 

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