Genus: Spinosaurus STROMER, 1915
Etymology: Latin, spina, “spine” and Greek sauros, “lizard”, Spine lizard.

Species: aegyptiacus STROMER, 1915
Etymology: In reference to the country it was found, Egypt.

Holotype: IPHG 1912 VIII 19

Locality: 3 km north of the Gebel el Dist in the basin of Baharija Oasis, Egypt.

Horizon: Baharija Formation.

Biostratigraphy:

Age: ?Early Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowemrost Gulf Epoch, Middle Cretaceous.

Material: Dentary with teeth, cervical, dorsal, sacral and caudal vertebrae, ribs.

Two verisons of a short skull.

Two versions of a longer skull. The bottom is after a new undescribed skull.

Three possible skeletal reconstructions.

Possible bending of the back (George Olshevsky).

Referred material:

MILNER, 1996, 2001

Locality: Taouz, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Material:

BMNH R 16420: Premaxilla, and fragmentary maxilla.

BMNH R 16421: Left dentary.

 

= Spinosaurus cf. aegyptiacus BUFFETAUT, 1989

Locality: Near the town of Taouz, at the base of a vast plateau known as the Hammada du Guir, Morocco.

Horizon: Continental Red Beds.

Biostratigraphy:

Age: Upper Albian Stage, Middle Gallic Subepoch, Uppermost Early Cretaceous Epoch or Lower Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

IMGP 969-3: Tooth.

IMGP 969-1: Fragmentary right maxilla.

 

= Spinosaurus cf. aegyptiacus BUFFETAUT & OUAJA, 2002

Locality: Jebel Miteur, Tataouine Governorate, Southern Tunisia.

Horizon: Chenini Sandstones.

Biostratigraphy:

Age: Early Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

ONM BM 231: Rostral part of left dentary containing 4 alveoli.

 

= Spinosaurus cf. aegyptiacus DAL SASSO, MAGANUCO, BUFFETAUT & MENDEZ, 2005 (published in 2006)

Locality: Kem Kem, east of the town of Taouz, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

MSNM V4047: Snout.

Note: Has an isolated vertebra of ?Onchopristis sp embedded between the right second premaxilary alveolus and its erupting tooth.

 

Locality: Kem Kem beds, N30°02', W5°12', near the outpost in Keneg ed Dal, Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

UCPC-2: Pair of fused nasals.

AZUMA, 2005

FPDM-V7011: Cast of a new undescribed skull.

Number: Not given: Dorsal vertebra.

IBRAHIM, SERENO, DAL SASSO, MAGANUCO, FABBRI, MARTILL, ZOUHRI, MYHRVOLD, & IURINO, 2014

UCRC PV601: Anterior dorsal vertebra.

FSAC-KK 11888: Neural spine, left ilium, right manual II-1 phalanx, left femur, right tibia, and right pedal ungual..

CA30: Mid cuadla vertebrta.

UCRC VP5: Anterior caudal vertebra.

 

 

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Species: maroccanus RUSSELL, 1996
Etymology: In reference to the “roysume du Maroc” (kingdom of Morocco).

Holotype: NMC 50791

Locality: Taout, ‘Gres rouges infracenomaniens’, Tafilalt, Morocco.

Horizon: Tegane Formation.

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material: Median cervical vertebrae.

Referred material:

NMC 50832, 50833: Dentary fragments.

NMC 41768, 50790, 50791: Median cervical vertebrae.

NMC 50813: Dorsal neural arch.


D'ANASTASIO & CAPASSO, 2004

Del Museo di Storia delle Scienze Biomediche dell'Universita degli Studi "G. d'Annunzion, PP #481: Cervical vertebra with Osteoarthritis.

 

TAQUET & RUSSELL, 1998

Locality: Gara Samani, Tademait, Algeria.

Horizon:

Biostratigraphy:

Age: Albian Stage, Middle Gallic Subepoch, Upper Early Cretaceous Epoch, Early Cretaceous.

Material:

MNHN SAM 124: Rostrum containing both premaxillae, maxillae, vomers and fragment of midshaft of right dentary.

MNHN SAM 125: Fragment of premaxilla.

MNHN SAM 126-127: Centra of 2 cervical vertebrae.

MNHN SAM 128: Neural arch of a dorsal vertebra.

 


= S. cf aegypticus SERENO, DUTHEIL, IAROCHENE, LARSSON, LYON, MAGWENE, SIDOR, VARRICCHIO & WILSON, 1996

Locality: Kem Kem region of southeastern Morocco.

Horizon: Kem Kem Beds.

Biostratigraphy:

Age: Cenomanian Stage, Uppermiddle Gallic Subepoch, Lowermost Gulf Epoch, Middle Cretaceous.

Material:

Ministere de l’Energie et des Mines, Rabat, Morocco: Jaw fragments and teeth.
Note: May belong to a new genus.

ANONYMOUS, 2004

Number: Not given: Dentary.