Genus: Ceratosaurus MARSH, 1884
Etymology: Greek kerat (keras) “horn” and Greek sauros for lizard.

Species: meriani (GREPPIN, 1870) (nomen dubium)
Etymology:
= Ceratosaurus sp (as per MADSEN & WELLES, 2000)
= Megalosaurus meriani GREPPIN, 1870 (nomen dubium)
= Antrodemus meriani (GREPPIN, 1870) STEEL, 1970 (nomen dubium)
= Labrosaurus meriani (GREPPIN, 1870) JANENSCH, 1920 (nomen dubium)
= Allosaurus meriani (GREPPIN, 1870) OLSEHVSKY, 1991 (nomen dubium)

Holotype: MH 350

Locality: Moutier , Jura Bernois, near Moutier, Savoy, Kanton Bern, Switzerland.

Horizon: Unter-Virgula-Schichten.

Biostratigraphy:

Age: Portlandian, Middle Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material: Teeth, two dorsal vertebrae and a Panzerplatte.
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Species: nasicornis MARSH, 1884
Etymology: Latin, nasus ‘nose’, and Latin, cornice, ‘ornamental molding along a wall’; in reference to the ornamental nose horn.

Holotype: USNM 4735

Locality: Marsh-Felch’s YPM Quarry 1, Garden Park, Sec. 28, T17S, R70W, Canyon City, Fremont County, Colorado.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Skull, lower jaws, atlas, axis, 7 cervical vertebrae, 12 fragmentary dorsal vertebrae, complete sacrum, 50 caudal vertebrae, 20 chevrons, last thoracic rib, fragmentary cervical vertebrae, left scapula, left ulna, left radius, 4 left metacarpals, 3 left phalanges, 3 right metacarpals, 1 right phalanages, ilia, pubes lacking distal ends, ischia lacking distal ends, femora, right tibia, right fibula, right astragalus, right calcaneum, 1 tarsal bones of distal left row, left metatarsals, right metatarsals, 1 terminal left phalanx of digit IV, several skin ossicles.

Note: Left ulna has a pathology (pers. observation), left metatarsals pathological.

Note: One manus claw belongs to Allosaurus.

Referred material:

McINTOSH, 1981

CM 21706: Incomplete dentary.

 

= Ceratosaurus? nasicornis BRITT, 1991

Locality: Dry Mesa Quarry 1, sec. 23, T.50N, R.14, Mesa County, on the Uncompahgre Plateau, in Western Colorado.

Horizon: Lower Upper Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Lower Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

BYUVP 4853: Anterior caudal.

BYUVP 4951, 4952: Fragmentary post dorsal vertebrae.

BYUVP 5008: Fragmentary metatarsal III.

BYUVP 5010: Left metatarsal III.

BYUVP 4838, 4908, 5092, 5254, 8910, 8937, 8938, 8982, 9108, 9136, 9141, 9152, 9161, 9162: Caudal vertebrae.

BYUVP 8907: Dorsal vertebra.

BYUVP 8974: Posterior caudal.

BYUVP 9142, 9143, 9144: Fragmentary mid dorsal or post vertebrae.

BYUVP 9163: Very post caudal.

 


BRITT, 1991

Locality: Sand Beach Site, 25 km (15 miles) southeast of the town of Moore and 1.6 km (1 mile) north of interstate 70, San Rafael Swell, Utah.

Horizon: Salt Wash Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material:

BYUVP 12893: Uncrushed partially articulated skull, presacral vertebrae 13-20, a femur, pubes, and ischia.

 


BRITT, MILES, CLOWARD & MADSEN, 1999

Locality: Bone Cabin Quarry West, Albany County, Wyoming.

Horizon: Member? Morrison Formation.

Biostratigraphy: Zone ?

Age:Age?, Stage?, Epoch?, Late Jurassic.

Material:

NAMAL 2002-02-28: Fragmentary skull and fragmentary skeleton of a juvenile.

 

BRITT, SCHEETZ, STATDMAN & CHURE, 2003

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

BUY 17550: Fragmentary skeleton with ossified tissue, tendon bundles.

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Species: sulcatus (MARSH, 1896) emend MADSEN & WELLES, 2000
Note: Ceratosaurus sp (as per MADSEN & WELLES, 2000)
= Labrosaurus sulcatus MARSH, 1896
Etymology:

Holotype: YPM 1936

Locality: Reeds YPM Quarry 9, (Mammal Quarry), T22N, R77W, Sec. 12, Como Bluff, Albany County, Wyoming.

Horizon: Upper or Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: ?Premaxillary tooth crown or anterior dentary tooth.

Note: Was believed to belong to the following, but was later shown to be from Quarry 9 by MADSEN & WELLES, 2000

Locality: Nail Quarry, Como Bluff, Como Bluff East (East of Marshall road), Albany County, Wyoming.

Horizon: Lower Part, Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 2.

Age: Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

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Species: roechlingi JANENSCH, 1925 (nomen dubium)
Etymology:

Lectotype: MB R 2162

Locality: Mw, Tendaguru hill, Tendaguru, Mtwara, Tanzania, East Africa.

Horizon: Upper Dinosaur Member, Tendaguru Formation.

Biostratigraphy:

Age: Late Tithonian Stage, Upper Malm Epoch, Late Jurassic.

Material: Partial middle caudal vertebral centrum.

Referred material:

MB R 1934 (MW 5): Posterior caudal vertebra.

MB R 1935 (Mw 3): Anterior caudal vertebra.

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Species: ? stechowi (JANENSCH, 1920) RAUHUT, 2011 (nomen dubium)
= Labrosaurus stechowi JANENSCH, 1920 (nomen dubium)
Etymology:
= Antrodemus stechowi (JANENSCH, 1920) (nomen dubium)
= Antrodemus steschowi CHABLI, 1985 (sic)

Holotype: MB R 1083

Locality: Mahimbwi Valley, Om, Tendaguru, Mtwara, Tanzania, East Africa.

Horizon: Middle Dinosaur Member, Tendaguru Formation.

Biostratigraphy:

Age: Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Tooth.

Referred material:

MB R 1086-1093: 8 teeth.

 

Locality: Bolachikombe-tales, Tendaguru, Mtwara, Tanzania, East Africa.

Horizon: Tendaguru Formation.

Biostratigraphy:

Age: Late Jurassic.

Material:

Type c:

 


Locality: Stegosaurier-Graben (St), Tendaguru, Mtwara, Tanzania, East Africa.

Horizon: Tendaguru Formation.

Biostratigraphy:

Age: Late Jurassic.

Material:

Type e: Tooth.

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Species: willisobrienorum PICKERING, 1995
Etymology:
= Ceratosaurus magnicornis MADSEN & WELLES, 2000
Etymology: In reference to the proportionately larger nasal horncore.

Holotype: MWC 1

Locality: Callison’s Quarries, Fruita Paleontological area, Fruita, SW 1/4 , sec. 24, T1N, R3W, Ute Meridian, near Grand Junction, Mesa County, Colorado.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 4.

Age: Upper Part, Comobluffian age, Tithonian Stage, Uppermost Malm Epoch, Late Jurassic.

Material: Skull and nearly complete skeleton.

Breakdown as follows;

MWC 1.1.14 (old PF-QB-24I): Left premaxilla, (old PF-QB-24J) right premaxilla, (old PF-QB-24L), premaxilla (?), fragment & tooth.

MWC 1.1.8 (old PF-QB-24A): Left maxilla.

MWC 1.1.13 (old PF-QB-24K): Right maxilla.

MWC 1.1.3 (old PF-QB-24C): Nasals.

MWC 1.1.1 (old PF-QB-24E): Frontals.

MWC 1.1.2 (old PF-QB-24B): Left lachrymal.

MWC 1.1.40 (old PF-QB-24M): Right squamosal, (old PF-QB-24T) left squamosal.

MWC 1.1.10 (old PF-QB-24F): Left jugal.

MWC 1.1 (old PF-QB-24H): Right ? jugal, (old PF-QB-36B) 5th cervical vertebra, (old PF-QB-43) 6th cervical vertebra, (old PF-QB-12) 7th cervical vertebra, (old PF-QB-4) 8th cervical vertebra, (old PF-QB-1A) 9th cervical vertebra, (old PF-QB-1B) 1st pectoral vertebra, (old PF-QB-1C) 2nd pectoral vertebra, (old PF-QB-2) 2nd dorsal vertebra, (old PF-QB-44) 3rd dorsal vertebra, (old PF-QB-3A) 5th dorsal vertebra, (old PF-QB-3B) 6th dorsal vertebra, (old PF-QB-3C) 7th dorsal vertebra, (old PF-QB-14A) 8th dorsal vertebra, (old PF-QB-34) posterior dorsal vertebra, (old PF-QB-8) anterior caudal, (old PF-QB-14B) median caudal, (old PF-QB-16, 17, 30, 35) posterior caudal vertebrae.

MWC 1.1.5 (old PF-QB-24O): Left paraoccipital.

MWC 1.1.1 (old PF-QB-24P): Left postorbital, (old PF-QB-24R) premaxillary? Tooth, (old PF-QB-24S) maxillary tooth.

MWC 1.1.32 (old PF-QB-24Q): Cervical rib, (old PF-QB-36B) medium chevron.

MWC 1.1.15 (old PF-QB-33A), 1.1.16 (old PF-QB-33B), 1.1.17 (old PF-QB-33C), 1.1.18 (old PF-QB-33D): Dermal ossicles.

MWC 1.1.30 (old PF-QB-46): Dermal ossicle, (old PF-QB-21) left humerus, (old PF-QB-19) manual ungual, (old PF-QB-25) left femur, (old PF-QB-31) right femur, (old PF-QB-42) left tibia, (old PF-QB-26A) right tibia, (old PF-QB-26B) right astragalus, (old PF-QB-26C) right calcaneum, (old PF-QB-27) left metatarsal-II, (old PF-QB-37) distal end of a right metatarsal-II, (old PF-QB-6) metatarsal-III-1 (?),(old PF-QB-47) left metatarsal-III, (old PF-QB-28) left D-II-1 of the pes, (old PF-QB-38) right D-II-1 of the pes, (old PF-QB-32) left D-III-2 (?) of the pes (?), (old PF-QB-7) left D-IV-1 of the pes, (old PF-QB-45) right D-IV-1 of the pes, (old PF-QB-40) right D-IV-1 (?) of the pes.
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Species: dentisulcatus MADSEN & WELLES, 2000
Etymology: In reference to the persistent, parallel grooves on the medial surface of the premaxillary teeth and the anterior 3 teeth in the dentary.
= Species: Nova MADSEN & STOKES, 1963/STEEL, 1970

Holotype: UMNH 5278

Locality: Cleveland-Lloyd Dinosaur Quarry, approximately 20 miles south of Price, SE 1/4, SE 1/4, Sec. 21, T17S, R11E, SLBM. lat 39°19’00", Long. 110°41’15"W, San Rafael Swell Area, Carbon County, Utah.

Horizon: Brushy Basin Member, Morrison Formation.

Biostratigraphy: Zone 3.

Age: Upper Part, Comobluffian age, Kimmeridgian Stage, uppermost Malm Epoch, Late Jurassic.

Material: Premaxillae, right maxilla, the right quadrate, right quadratojugal, right coracoid, right scapula, elements from the fore limb and foot, and left Ilium.
Note: 1 1/2 times larger than the type.

Referred material:

UUVP 25, 80, 93, 102, 113, 208, 236, 351, 367, 433, 469, 507, 677, 6788: Dermal ossicles.

UUVP 47: Manual phalanx?

UUVP 48: 9th dorsal vertebra.

UUVP 49: Pectoral (?) vertebra.

UUVP 56: Left femur, right fibula.

UUVP 66: 3rd pectoral vertebra.

UUVP 81: Anterior caudal centra (sectioned).

UUVP 155: Right dentary.

UUVP 156: Right splenial.

UUVP 157: Left splenial.

UUVP 158: Left dentary.
Note: Pathology, the right dentary has a diastma that may be pathologic like “Labrosaurus ferox”.

UUVP 170: Anterior caudal vertebra (sectioned).

UUVP 173: Left fibula.

UUVP 234, 6949: Medial chevron.

UUVP 272, 6941: anterior chevron.

UUVP 316: Right dorsal rib.

UUVP 317: Right scapulocoracoid.

UUVP 327: Tarsal IV.

UUVP 329: 1st pectoral vertebra.

UUVP 365: Neural arch of an anterior caudal.

UUVP 375-376: 2 medial vertebrae.

UUVP 377-378: Distal chevrons of UUVP 375-376.

UUVP 531, 533, 6948, 6945-46: Distal chevrons.

UUVP 534, 532, 530, 212, 6951-56: Distal caudal vertebrae.

UUVP 548: Right anterior dorsal rib.

UUVP 549: Left humerus.

UUVP 562: 9th cervical vertebra.

UUVP 610: Left anterior dorsal rib.

UUVP 674: Left and right premaxillae, left maxilla.

UUVP 1035: 8th cervical vertebra.

UUVP 1053: Atlantal intercentrum, atlas/odontoid/axial intercentrum/axis, third cervical.

UUVP 1231: 7th dorsal.

UUVP 1229: 5th dorsal vertebra.

UUVP 1615: 7th cervical centrum.

UUVP 1646: Right pterygoid, right quadrate/quadratojugal, right jugal.

UUVP 1979: Right metacarpal II.

UUVP 2172: 5th left cervical rib.

UUVP 6520: 4th right cervical ribs.

UUVP 3729: Pedal phalanges.

UUVP 5681: Left tibia, left astragalus and left calcaneum.

UUVP 5682: Right tibia, right humerus, right astragalus.

UUVP 5683: Left metatarsal IV.

UUVP 5911: Right angular.

UUVP 5959: Distal caudal vertebra.

UUVP 5960: Medial caudal with chevron.

UUVP 6938: 8th dorsal centrum.

UUVP 6964: 6th cervical vertebra.

UUVP 6695: 5th cervical vertebra.

UUVP 6743: Left and right premaxillae.

UUVP 6958: Distal caudal with chevron (6957).

UUVP 6961, 6305, 53 (6939) 438, 439: Anterior caudal vertebra.

UUVP 6963: 4th cervical vertebra.

UUVP 6964: 6th cervical vertebra.

UUVP 6965: 5th cervical vertebra.