Genus: Styracosaurus LAMBE, 1913
Etymology: Greek, styrak- (styraz) “spike on the end of spear” and Greek, sauros, “lizard”: “spike lizard”.
= Stylacosaurus CHARIG & HORSFIELD, 1975 (sic)
= Rubeosaurus McDONALD & HORNER, 2010
= Genus: Nova McDONALD & HORNER, 2007
Etymology: Latin, rubes, adjective "thornbush or bramble" and Latinized Greek noun, saurus, 'lizard'; Thornbush lizard, in reference to the array of spikes adorning the parietals.

Species: albertensis LAMBE, 1913
Etymology: In reference to the province of Alberta in Canada.
= Monoclonius albertensis (LAMBE, 1913) LEAHY, 1987

Holotype: CMN (NMC) 344

Locality: Quarry 016, Dinosaur Provincial Park, about 1 1/2 miles southeast of Denhart Ferry, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material: Skull and skeleton.
Note: A year later after the find of the skull, the skeleton was discovered.
Note: Pathological parietal (HOLMES, RYAN & LLYOD, 2007, TANKE & ROTHSCHILD, 2010)

After RYAN, HOLMES, & RUSSELL, 2007

Referred material:

BROWN, HOLMES & CURRIE, 2020

Locality: Princess Coule, jsut south of Dinosaur Provincal Park, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 2009.080.0001: Skull.

 

CURRIE & RUSSELL, 2005

Locality: Quarry 124, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

BM R8648: Fragmentary skull and skeleton.

 

Locality: Quarry 183, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 87.52.1: Fragmentary skull.

 

Locality: Quarry unknown, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

ROM 1426, 5361: Fragmentary skulls.

 

EVANS, McGARRITY, & RYAN, 2014

Locality: Sage Creek Grazing Reserved, approximately 10 km north of the Onefour Research Station, Cypress County, southern Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 1989.097.0001: Juvenile specimen.

 

HOLMES, PERSONS, RUUPAL, QURESHI & CURRIE, 2020

Locality: Quarry Q262, along Matzhiwin Creek near Duchess, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

UALVP 55900: Skull and lower jaw.

 

RYAN, 2003, HOLMES, RYAN & MURRAY, 2006

Locality: Styracosaurus bone bed (mentioned by CURRIE & DODSON, 1984), bone bed 42 (BB 42), Dinosaur Provincial Park, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 66.10.3: Parietal P3 & 4.

RTMP 66.10.4, 81.19.160, 81.19.248, 90.58.5, 90.58.5, 90.58.7,92.36.254, 81.19.209: post parietal bars.

RTMP 66.10.6: Right lateral parietal.

RTMP 66.10.10-13, 66.10.5, 81.19.211, 81.19.223, 82.19.54, 83.36.158, 98.68.33, 66.10.9: Parietal spikes.

RTMP 66.10.14, 81.16.360, 18.191.141, 91.19.215, 91.19.223, 90.58.2, 90.58.9, 92.12.34, 99.55.224: Parietal fragment.

RTMP 66.10.15: Parietal P3? base.

RTMP 66.10.16: Parietal P3 base.

RTMP 66.10.17, 24, 40, 79.13.4, 90.58.10, 90.58.3,90.58.6, 91.58.12, 98.93.42, 90.58.15: Fragmentary nasals.

RTMP 66.10.19-23, 94.12.886, 81.19.156: Nasals.

RTMP 66.10.26: Left dentary fragment.

RTMP 66.10.27, 2001.12.10: Left dentary.

RTMP 99.55.2: Fragmentary parietal.

RTMP 66.10.28, 66.20.32, 93.36.166, 97.12.122, 99.55.3-5, 2001.12.4: Parietals.

RTMP 66.10.31, 81.19.282: Right jugals.

RTMP 66.10.33, 81.19.214, 81.19.266, 99.55.6, 2001.12.5: Left squamosals.

RTMP 66.10.34, 66.10.29, 2001.12.3, 2001.12.7, 2001.12.8, 2001.12.9, 66.10.30: Right squamosals.

RTMP 66.10.37: Sacrum.

RTMP 66.10.39: Right postorbital.

RTMP 66.10.41, 90.58.4: Left supraorbital.

RTMP 66.10.42, 81.19.216: Rostral fragment.

RTMP 66.10.43: Predentary.

RTMP 67.8.15: Left frontal.

RTMP 79.13.3: Cervical centrum.

RMTP 81.19.137: Preoximal caudal vertebra.

RTMP 81.19.138: Metatarsal.

RTMP 81.19.139: Left? quadrate.

RTMP 81.58.155: Nasal.RTMP 81.19.158: Ischium.

RTMP 81.19.208: Rib fragment.

RTMP 81.19.210: Occipital condyle.

RTMP 81.19.213: Squamosal.

RTMP 81.19.217: Skull element.

RTMP 81.19.222: Left lateral parietal.

RTMP 81.19.247, 94.12.896: Premaxillae.

RTMP 81.58.155: Nasal.

RTMP 83.36.154:Parietal with spikes.

RTMP 88.36.26, 90.58.14: Quadratojugal fragments.

RTMP 90.58.8, Supraorbital.

RTMP 90.58.17, DHT/DPP-157: Laterial parietals.

RTMP 90.58.18: Right jugal and epijugal.

RTMP 94.12.394: Base of spikes 3 and 4.

RTMP 94.12.699: Right angular.

RTMP 94.12.890: Orbital horncore.

RTMP 94.12.894: Predentary fragment.

RTMP 94.12.899, 90.58.11: Spike fragment.

RTMP 98.93.39: Angular.

RTMP 98.93.43: Nasal and rostral fragments.

RTMP 98.93.64: Right supraorbital.

RTMP 99.55.3: Right coracoid fragment.

RTMP 81.19.160, 91.36.254, 99.55.5: Partial parietals.

RTMP 99.93.41: Parietal P1.

RTMP 1999.55.1: Postorbital.

RYAN, HOLMES & RUSELL, 2007

RTMP 88.36.20: Large adult partial parietal.

RTMP 99.52.2: Fragmentary parietal.

TANKE & ROTHSCHILD, 2010

RMTP 90.58.1: Dorsal neural arch with coarse pitting on the right postzygapophysis, possibly related to spondyloarthropathy.

RTMP 89.97.1: Nearly complete tail with caudals 27-30 with cheverons thoroughly fused and for a swollen mass.

 

Locality: BB20, Dinosaur Provincial Park, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 98.93.133: Left supraorbital.

 

 

Locality: BB55, Dinosaur Provincial Park, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 98.03.163: Right nasal.

RTMP 98.93.134, 98.93.154, 98.93.38: Parietal fragments

TANKE, 2007

RTMP 98.93.163: Juveinile nasal horncore (cf. Styracosaurus)

 

Locality: Sandy Point, Dinosaur Provincial Park, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 90.11.4: Parietal.

RTMP 90.117.4: Laterial spike.

 

RYAN, HOLMES, & RUSSELL, 2007

Locality: West branch of Sand Creek, Red Deer River Valley, about 2 miles south west of the mouth of the creek and about 200 feet above the river, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

ROM: 1436: Partial parietal.

 

SWEENEY & BOYDEN, 1993

Locality: Along the Milk River, just south of the 49th parallel, Montana.

Horizon: Judith River Formation.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Right parietal, predentary, nose horn, left femur, left ilium, base of occipital, right humerus, a dorsal vertebra and 5 ribs.

 

TANKE PERS. COM., RYAN, HOLMES & RUSSELL, 2007

Locality: Quarry 179, Dinosaur Provincial Park, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 86.126.1: Skull missing nasal horn of a juvenile.
Note: Was thought to be a Centrosaurus.

RYAN, EBERTH, BRINKMAN & RUSSELL, 1998

RTMP 89.36.20: Fragmentary skull.

RTMP. 89.97.1: Fragmentary skull.

TANKE, 2007

RTMP 86.126.1: Skull missing both nasals and premaxillae and no maxillary teeth of a subadult.

 

TANKE, 2007

Locality: 100 meters northwest of Quarry 016, Dinosaur Provincial Park, about 1 1/2 miles southeast of Denhart Ferry, Dinosaur Provincial Park, Red Deer River, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 98.93.64: Right orbital region with sutures.

= Styracosaurus parksi BROWN & SCHLAIKJER, 1937
= Styracosaurus borealis BROWN via OLSHEVSKY, 1992
Note: The species name was never published but was on a file photograph of Styracosaurus parksi at the AMNH.
Etymology: In honor of William Arthur Parks.
= Monoclonius sp LULL, 1933

Holotype: AMNH 5372

Locality: Quarry unknown (Quarry 250 ?), Dinosaur Provincial Park, middle fork of Sand Creek on the right bank, approximately 250 feet above the Red Deer River, 12 miles below Steveville, Newell County, Alberta Province, Canada.

Horizon: Upper Dinosaur Park Formation, upper Judith River Group.

Biostratigraphy:

Age: Judithian age, Middle Campanian Stage, middle Senonian subepoch, upper Gulf Epoch, Late Cretaceous.

Material: Fragmentary skull, fragmentary dentary and a nearly complete skeleton.


_____________________________________________________________________________________

Species: ovatus GILMORE, 1930
= Rubeosaurus ovatus (GILMORE, 1930) McDONALD & HORNER, 2010
Etymology: Latin, ovatus, "oval."
= Taxon A* Similar to Styracosaurus HORNER, VARRICCHIO & GOODWIN, 1992

Holotype: USNM 11869

Locality: T37N, R8W, Milk River, Blackfeet Indian Reservation, Glacier County, Montana.

Horizon: Two Medicine Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material: Posterior portion of frill and numerous fragments.

 

Referred material:

GILMORE, 1939

Locality: South Side of Milk River, Vicinity of Landslide Butte on Blackfeet Nation, Glacer County, Montana.

Horizon: Two Medicine Formation.

Biostratigraphy:

Age: Judithian age, Campanian Stage, Senonian subepoch, Upper Gulf epoch, Late Cretaceous.

Material:

USNM 14765: Right half of frill lacking the squamosal, articulated left postorbital, jugal, postfrontal and prefrontal, lachrymal and supraorbital, right maxillary containing several teeth, left premaxillary, rostal, a portion of the right half of the nasal horn core, fragment of the basioccipital, unidentified skull fragments, both femora, left scapula, first rib, one dorsal vertebra, and two phalangeial bones. An adult.