Genus: Protoceratops GRANGER & GREGORY, 1923
Etymology: Greek, protos, “first” and Greek, ceratops, “horn face”: “first horn face”.
= Proceratops DONG, 1979 non LULL, 1906 (sic)

Species: andrewsi GRANGER & GREGORY, 1923
Etymology: In honor of Roy Chapman Andrews.
= Protoceratops andrewis [ANONYMOUS] 1990 (sic)

Holotype: AMNH 6251

Locality: Kweiwa-ting trail, east of Artsa Bogdo, Lat 44’N, Long 104’E, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Skull lacking the occiput and tip of premaxilla.

Protoceratops andrewsi (modified from GRANGER & GREGORY, 1923), Holotype: AMNH 6251.

Referred material:

BROWN & SCHLAIKJER, 1940

Locality: Bayan Zag (= Bayn Dzak, Shabarakh Usu, Flaming Cliffs), District of Dalandzadgad, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Male skeleton walking on all fours.

Male walking bipedally.

Male skull.

Pes and manus showing the flat phalanges used for digging.

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Female skull.

Juvenile skull.

AMNH 6273: Skull and jaws of a juvenile.

AMNH 6274: Miscellaneous jaw fragments and foot-bones.

AMNH 6409: Skull, nearly perfect.

AMNH 6416: Skull, jaws and skeleton, complete to proximal caudals.

AMNH 6417: Skull jaws and nearly complete skeleton.

AMNH 6418: Skull, jaws and most of skeleton to middle to tail.

AMNH 6419: Skull, jaws, portions of anterior part of skeleton of a juvenile.

AMNH 6421: Skull and jaws in concretion of a juvenile.

AMNH 6422: Fragmentary skull and jaws, some skeletal fragments of a juvenile.

AMNH 6423: Skull and jaws in concretion, weathered.

AMNH 6424: Large part of skeleton, without skull and jaws.

AMNH 6425: Skull and jaws, nearly perfect.

AMNH 6426, 6450: Skull, jaws and part of skeleton.

AMNH 6429: Skull, not teeth.

AMNH 6408, 6413, 6414, 6428, 6430, 6431, 6432, 6434, 6439, 6444, 6447, 6448, 6461, 6489, 6491: Skulls and jaws.

Protoceratops andrewsi (after LULL, 1933), AMNH 6408.

AMNH 6433: Skull and jaws, fine.

AMNH 6436: Teeth.

AMNH 6437: Skull and jaws in concretion.

AMNH 6438: Skull and jaws, large.

AMNH 6440, 6441: Skulls and jaws, panel mount.

AMNH 6442: Imperfect skull.

AMNH 6443: Skull and jaws, crushed.

AMNH 6445, 6446: Fragmentary skulls in matrix.

AMNH 6449, 6458, 6458: Fragmentary skulls.

AMNH 6451: Fragmentary skull and part of skeleton.

AMNH 6453: Fragmentary skeleton.

AMNH 6454: Weathered skull and much of skeleton.

AMNH 6459, 6460, 6463: Lower jaws.

AMNH 6465: Skull and partial skeleton.

AMNH 6466: Large skull and jaws, and thoracic section.

AMNH 6467: Skull, jaws, and nearly complete skeleton, mounted.

AMNH 6459: Skull fragments and parts of skeleton.

AMNH 6470: Back of skull and parts of skeleton.

AMNH 6472: Incomplete skull and jaws.

AMNH 6473: Part of skull and jaws, poor.

AMNH 6474: Caudal vertebrae and hind limbs.

AMNH 6475: Skeletal parts.

AMNH 6477: Imperfect skull.

AMNH 6478: Caudals, ulna, radius, fore and hind foot.

AMNH 6479: Partial skeleton.

AMNH 6480: Partial skull and jaws in weathered concretion.

AMNH 6481: Pelvis.

AMNH 6482: Fore and hind foot.

AMNH 6483: Weathered skull and jaws, fragmentary and small.

AMNH 6484: Skeletal parts.

AMNH 6485: Skull in weathered concretion.

AMNH 6486, 6487: Weathered skulls and jaws.

AMNH 6488: Fragmentary skull in weathered concretion.

AMNH 6490: Part of skeleton.

AMNH 6492: Fragmentary skeleton.

AMNH 6493: Vertebral column.

AMNH 6494: Pelvis, hind limb and series of vertebrae.

AMNH 6495, 6496: Front of skulls and jaws.

AMNH 6497: Fragment of lower jaw.

AMNH 6498: Left dentary of a very young individual.

AMNH 6499: Right dentary of an unborn.

AMNH 6636: Poor skull and part of skeleton.

AMNH 6637: Part of a skull.

AMNH 6638: Front of skull.

AMNH 6639: Portion of a skull.

AMNH 6640: Caudal vertebrae.

AMNH 6644: Parts of small skull and jaw, limb-bones, etc.

AMNH 6415: Skull and jaws, small and crushed.
Note: Sent to Peking, China, 1925.

AMNH 6420: Skull and jaws, largest size.
Note: Sent to Field Museum, Chicago, Jan. 1926.

AMNH 6427: Skull and jaws.
Note: Sent to Univ. California, 1934.

AMNH 6435: Skull, jaws, most of skeleton.
Note: Sent to Field Museum, Chicago, Jan. 1926.

AMNH 6455: Hind foot.
Note: Sent to Munich, Germany, 1933.

AMNH 6456: Skull and jaws.
Note: Sent to Paris, France, 1927.

AMNH 6457: Half of a skull and jaw fragment.
Note: Sent to Munich, Germany, 1933.

AMNH 6452: Lower jaw, palate, foot bones.
Note: Sent to Uinv. California, 1934.

AMNH 6464: Skull, jaws, part of skeleton.
Note: Sent to Urga, Mongolia, 1927.

AMNH 6476: Skull and jaws.
Note: Sent to Urga, Mongolia, 1927.

AMNH 6641: Part of skull and jaws.
Note: Sent to Munich, Germany, 1933.

AMNH unnumbered: Front of skull and jaws.
Note: Sent to Public School Mus. in Battle Creek, Mich.

AMNH unnumbered: Posterior part of skull in concretion, weathered.
Note: Sent to Public School Mus. in Battle Creek, Mich.

The following eggs were referred to Protoceratops, but probably do not belong to this genus.

AMNH 2905: Portion of egg, small.

AMNH 6505: Portions of 2 eggs.

AMNH 6506: Cast of egg, with few fragments of shell adhering.

AMNH 6507: Finely preserved fragments of 1 or more eggshells.

AMNH 6508: Group of 15 or more or less complete eggs and fragments of probably 2 others which were weathered out and broken up. 13 in situ, 1 sent to Colgate University, 1924.

AMNH 6509: Group of 3 weathered eggs, 2 showing portion of embryos.

AMNH 6510: Group of 3 weathered eggs.

AMNH 6511: Group of 5 eggs in matrix.

AMNH 6512: Group of several eggs in matrix, each partly weathered off.

AMNH 6513: Group of 5 eggs in matrix.

AMNH 6631: Group of 18 eggs, each partly weathered off.

AMNH 6633: 8 large eggs, crushed but fairly complete, parts of several others.

AMNH 6635: Group of 6 small eggs.
Note: Sent to the Field Museum, Chicago, 1926.

AMNH 6642: Nest of 4 small, thin-shelled eggs.

AMNH 6649: Group of badly weathered eggs.

AMNH 6650: Egg.

AMNH 6651: Egg fragments.

AMNH 6652: Eggs-crushed.

AMNH 6654: Very small egg in concretion. Probably crocodilian?

AMNH 6660: Fragments of eggshells from 1 knoll, probably from 1 clutch.

Note: A furcula has been found with a skeleton.

FOSTOWICZ-FRELIK & SLOWIAK, 2018

ZPAL MgD-II/3a-d: Subadult skeleton.

ZPAL MgD-II/8: Adult skeleton.

ZPAL MgD-II/11a-c: Adult skeleton.

ZPAL MgD-II/15: Skeleton of a subadult.

ZPAL MgD-II/16a, b: Adult skeleton.

ZPAL MgD-II/33: Adult skeleton.

ZPAL MgD-II/407, /408: Juvenile skeletons.

GIERLINGSKI, LOCKLEY, SINGER & NIEDZWIDZKI, 2008, 2011

ZPAL Mg D-II/3: Articulated skeleton.

Note: 1 pes print was found under the skeleton.

HOPSON, 1979

AMNH 6466: Braincase.

Note: CHURE, 2000 states that PIN 3247/7, 3148/8: Skulls are stolen.

KIELAN-JAWOROWSKA & BARSBOLD, 1972

Numbers: Not given: Skeleton, skull and several fragments of skeletons.

KURZANOV, 1972

PIN 614-6: Fragmentary skull of an adult female.

PIN 614-9, 614-15: Fragmentary skulls.

McINTOSH, 1981

CM 9185: Fragmentary skull, fragmentary skeleton.

CM 9186: Hyoid of the above specimen.

SANEYOSHI, WATABE, TSUBAMOTO, TSOGTBAATAR, CHINZORIG & SUZUKI, 2010

Field number: 070619 BD KWR:

SUZUKI & WATABE, 2000b

Field number 980826 BDz Proto. COY: Skull.

TERESHCHENO, 1992/1994

PIN 614-41/1: Distal half of a femur.

PIN 614-45/1: Proximal half of a large tibia.

PIN 614-53/1: Pelvic girdle articulated with the right hind limb.

PIN 614/601: A skull.

TERESCHCHENO, 2008

PIN 614/61: Fragmentary skeleton of a young adult female.

PIN 614/31: Fragmentary skeleton of a young adult female.

PIN 614/35: Fragmentary skeleton of a young adult male.

PIN 614/30: Fragmentary skull and skeleton of an adult female.

ZPAL, no. MgD-II/4: Fragmentary skull and skeleton of an old male.

TSOGTBAATAR & CHINZORIG, 2010

Field number: 070619 BDz Proto-mtt. KWR: Nearly complete right hind limbwithout femur and part of the left pedial digits, distal end of the tibia and fibula and bone fragments.

 

BOHLIN, 1953

Locality: Ulan-tsonch, at the northern caravan route from eastern China to Sinkiang, Ordos (Shaan-Gan-Ning) Basin, Inner Mongolia (Nei Mongolia), China.

Horizon: Ulan-Tsonch Formation.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given: Teeth and centrum.

 

CARPENTER & ALF, 1994

Locality: Ikh Shoonkht, Ömnögov (South Gobi), Mongolia.

Biostratigraphy:

Age: Djadochta Formation, Late Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Skull and skeleton of a juvenile.

Protoceratops andrewsi (after CARPENTER & ALF, 1994).

Protoceratops andrewsi (after CARPENTER & ALF, 1994).

 

 

CZEPINSKI, 2020

Locality: Zamyn Khnod locality, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

MPC-D 100/505: A weathered skull.

 

Locality: Zhinst Tologi site, Üüden Sair Area, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

MPC-D 100/551: A nearly complete articulaed skeleton with skull and mandible.

 

DONG, 1992

Locality: Bayan Mandahu, Urad Houqi, Inner Mongolia (Nei Mongolia), China.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: 28 skulls of embryos, juveniles and adults and some with skeletons.

DONG & CURRIE, 1993

IVPP 9606: Nearly complete skull of an embryo.

 

FASTOVSKY, BADAMGARAV, ISHIMOTO, WATABE & WEISHAMPEL, 1994, 1996

Locality: Tögrögiin Shiree (= Tugrugeen Shireh, Tugrikin-Shireh locality), 44°13’N, 103°16’E, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Numbers: Not given: Skull, snout upward, resting on occiput, snout obliterated by weathering.

Numbers: Not given: Scapula, isolated, lying along bedding plane.

Numbers: Not given: Skull and isolated, missing frill and upper part, lower jaw.

Numbers: Not given: Post-crania, complete skeleton from scapula back to tip of tail lying on right half of body, anterior opposite dip direction of cross section.

Numbers: Not given: Skull flattened, disarticulated, mandible inverted over fragments of skull roof and frill.

Numbers: Not given: Pelvis and long bones disarticulated bones lying along bedding plane.

Numbers: Not given: Post-crania articulated trunk and pelvis with flexed real limbs, lying ventral down, anterior towards dip direction of cross section.

Numbers: Not given: Skull and postcrania articulated specimen minus forelimbs and tail, lying ventral down, anterior in dip direction of cross strata.

Numbers: Not given: Skull and postcrania articulated trunk with disarticulated limbs and frill material, anterior opposite dip direction of cross strata.

Numbers: Not given: Skull and postcrania, skull and neck, anterior towards dip direction of cross-strata.

Numbers: Not given: Skull and postcrania, disarticulated skull, pelvis, scapula, and ribs oriented relative to each other as in life, with ventral down, anterior opposite dip direction of cross-strata.

Numbers: Not given: Skull and mandible resting on mandible rami.

= Protoceratops cf. andrewsi FASTOVSKY, WEISHAMPEL, WATABE, BARSBOLD, TSOGTBAATAR & NARMANDAKH, 2011

MPC-D 100/530: Nest of 13 babies.

KIRKLAND, 1997, KIRKLAND & BADER, 2007

In situ: Fragmentary skull and skeleton with insect borings and pupae.

HONE, FARKE, WATABE, SHIGERU, & TSOGTBAATAR, 2014

MPC-D 100/526: Four juveinle individuals.

KIELAN-JAWOROWSKA & BARSBOLD, 1972

Number: Not given: 2 skulls and 2 skeletons.

Note: One skeleton is the one in mortal combat with Velociraptor, the other is a “standing” skeleton.

KURZANOV, 1972

PIN 3143/4-1: Skull of a adult male missing some of the frill and snout.

PIN 3143/1-1, 2-1. 3-1: Fragmentary skulls.

NIKOLOFF & HUENE, 1966

Number: Not given: 20 skeletons.

NOVACK, NORELL, McKENNA & CLARK, 1994

AMNH: 12 jumbled skeletons.

OSMOLSKA, 1986

GIN 100/512: Skull.

TERESHCHENO, 1992/1994

PIN 3143/7: Skull and skelton of a young adult female.

PIN 3143/5: Skull and skeleton of a young adult male.

SANEYOSHI, WATABE, SUZUKI & TSOGTBAATAR, 2011

MPC-D 100/533, /534: Fragmentary skulls and skeletons.

Note: With beetle borrings on the skull and skeletons.

SANEYOSHI, WATABE, TSUBAMOTO, TSOGTBAATAR, CHINZORIG & SUZUKI, 2010

Field number: 070619 TS TSUI: Juvenile specimen.

Field number: 070619 TS SZK: Lower jaw.

Field number: 070620 TS TSGT: Standing skeleton.

TERESCHCHENO, 2008

PIN 3143/6: Fragmentary skeleton of a juvenile female.

PIN 3143/12, 16: Fragmentary skeletons of adult male.

PIN 3143/14: Fragmentary skeleton of an adult female.

PIN 3143/9, 4: Fragmentary skeletons of old adult females.

WATABE & SUZUKI, 2000a

Field number 930828-29 TS-II-1 (HMNS 940302): Complete skeleton.

Field number 930830 TS-II-2, 930902 TS-I-1: Posterior half of skeletons.

Field number 930902 TS-I-2: Part of skull and lower jaw.

Field number 930902 TS-111: Skull.

 

FOSTOWICZ-FRELIK & SLOWIAK, 2018

Loocality: Toogreek, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

ZPAL MgD-II/35a-c: Subadult skeleton.

 

HU, CHENG, PANG & FANG, 2001

Locality: Vicinity of Zhangying, Jiaoxian County adjacent to Zhucheng County, China.

Horizon: Wangshi Group.

Biostratigraphy:

Age: Late Cretaceous.

Material:

Numbers: Not given: Maxilla, dentary, isolated teeth, humerus, ilium, femur, tibia, a number of vertebrae and some fragmentary bones.

Numbers: Not given: Scapula, rib, humerus, femur, fibula, sacrum and some vertebrae.

 

KIELAN-JAWOROWSKA & BARSBOLD, 1972

Locality: Khermeen Tsav, Mongolia.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Number: Not given:

 

 


NOVACK, NORELL, McKENNA & CLARK, 1994

Locality: Mongolia.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH: Less than nine centimeters long skeletons of newborns?

 

 

Locality: Khugene Tsavkhlant, Mongolia.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

AMNH:

 


SANEYOSHI, WATABE, TSUBAMOTO, TSOGTBAATAR, CHINZORIG & SUZUKI, 2010

Locality: Bortologi locality, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Field number: 070627 BGL TSGT: Skull and skeletons.

 

SUZUKI & WATABE, 2000b

Locality: Udyn Sayr, Nemegt Basin, Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Field number 98019 US Proto NAR: Skeleton.

Field number 980821 US Proto ENKH: Skull and skeleton.

Field number 980821 US SZK 9:05: Skull.
Note: Zhinst Tologi.

HANDA, WATABE, & TSOGTBAATAR, 2012

MPC-D 100-537: Skull with articulated mandible of a subadult.

= Protoceratops cf. andrewsi MPC-D 100/538: Skull with mandible of a subadult.

MPC-D 100/539: Skull without mandible, a cervical vertebra n dpartial pess, of a subadult.

SANEYOSHI, WATABE, TSUBAMOTO, TSOGTBAATAR, CHINZORIG & SUZUKI, 2010

Field number: 070626 US KWR: Fragmentary skull and skeleton.

MATSUMOTO, HASIHIMOTO, SONODA, FUJIYAMA, MIFUNE, KAWAHARA & SANEYOSHI, 2010

Field number 70626 US KWR Proto 1/3 (HMNS 2008-06-11): Part of fril.

Field number 70626 US KWR Proto 2/3 (HMNS 2008-06-11): Part of frill.

Field number 070627 US KWR Proto 3/3 (HMNS 2006-06-013): Part of frill.

 

 

TERESHCHENO, 1992/1994

Locality: Gil’bentu, Mongolia.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

PIN 4550/1: 3 first cervical vertebrae.

 

TSOGTBAATAR & CHINZORIG, 2010

Locality: Bor Tologi, South of Arts Bogd Mountian, Mongolia.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Field number: 070627 BLG Proto-skull TSGT: Skull without mandible.

 

WATABE & SUZUKI, 2000a

Locality: Tögrögiin Shiree II (= Tugrugeen Shireh, Tugrikin-Shireh II locality), Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Field number 930828-1 TS-II: Body parts.

Field number 930828-2 TS-II: Body parts.

WATABE & SUZUKI, 2000b

Field number 940728 TS-II GK: Skeleton.

 


WATABE & SUZUKI, 2000b

Locality: Tögrögiin Shiree-IV (= Tugrugeen Shireh, Tugrikin-Shireh-IV locality), Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Field number 940727 TS-IV GK, ISHI: Skeletons.

Field number 940727 TS-IV NAR: Small skull.

TSOGTBAATAR & CHINZORIG, 2010

Field number 940726 TS-IV Tsog: Nearly complete specimen.

 

 

 

Locality: Tögrögiin Shiree-I (= Tugrugeen Shireh, Tugrikin-Shireh-I locality), Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Field number 940801 TS-I SZK: Small skeleton.

 

WATABE, SUZUKI, TSOGTBAATAR, BARSBOLD & WEISHAMPEL, 1996

Locality: Tögrögiin Shiree-V (= Tugrugeen Shireh, Tugrikin-Shireh-V locality), Ömnögov (South Gobi), Mongolia.

Horizon: Djadochta Formation.

Biostratigraphy:

Age: Middle Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material:

Field number 940728 TS-V NAR: A nest of 15 juveniles.
Note: Maybe from a burrow.

WATABE & SUZUKI, 2000b

Field number 940725 TS-V MTS: Skeleton.

Field number 940728 TS-V MTS: Skeleton.

_____________________________________________________________________________________

Species: hellenikorhinus LAMBERT, GODEFROIT, LI, SHANG & DONG, 2001
Etymology: Greek, Hellenikos, "relative to ancient Greece" and Greek, rhins, "nose," with its elevated and angular snout, this species has some kind of a greek profile.

Holotype: IMM 95BM1/1

Locality: Quarry 95BM1/1, N41°44’485:, E106°44’703, Altitude 1307, Bayan Mandahu, Urad Houqi Banner, Bayan Nor League, Inner Mongolia, China.

Horicon: Bayan Mandahu Formation, ?correlated with the Djadokhta Formation.

Biostratigraphy:

Age: Campanian Stage, Senonian Subepoch, Gulf Epoch, Late Cretaceous.

Material: Nearly complete skull of a ‘male’.

Paratype:

IMM96BM1/4: Nearly complete skull of a ‘female’.

Referred material:

IMM 96BM1/1: Large Skull.

IMM 96BM5/5: Large skull.

IMM 96BM5/2: Right dentary, 4 teeth, right squamosal and quadrate.

IMM 96BM1/7: Large skull.

IMM 96BM2/1: Small skull.

IMM 96BM5/3: Partial very small skull.

IMM 96BM6/4: Left maxilla and jugal, right hind limb and series of caudal vertebrae.