Species: rex OSBORN, 1905
Etymology: Greek, rex, the king: “Tyrant lizard, the King”.

Holotype: CM 9380 (Formerly AMNH 973)

Locality: Near Jordan, Dawson County, Montana.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Partial skull and mandible, right? ectopterygoid misidentified as a right postorbital, one cervical, seven dorsals, five sacrals, three abdominal ribs, right scapula, left humerus, ilia, pubes, ischia, left femur and part of right, tibia, right metatarsals II, and III, left metatarsal IV of a robust individual.

Referred material:

AMNH 5027: Skull and jaws, cervical, dorsal, and most of caudal vertebrae, ribs, cervical ribs, ilium, pubis and ischium of a more gracile individual.
Note: Paleopathologies, dorsals 7 and 8 exhibit pathological fusion of the centra (NEWMAN, 1970), the centra of cervical 10 and dorsal 1 are also fused (ROTHSCHILD, 1997).

CM 9379 (Formerly AMNH 5029): Occiput and Brain case, sectioned.

 

 

Locality: On the Lester D. Engdahl Ranch (not, as originally noted, the F. S. McKeever Ranch), S34, T21N, R37E, near Jordan, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

LACM 23844: Disarticulated skull and fragmentary skeleton.
Note: Pathological openings in the right surangulars.

 

 

Locality: Glasgow, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BMNH R7995 (Formerly AMNH 5881): Femur, right and left tibiae, metatarsal I left, right and left metatarsal II, Metatarsal 1V, and abdominal ribs.

Note: Slightly smaller than the type.

 

Locality: Converse County, Wyoming

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 5117: Occiput, pterygoid, hyoid, and brain case of an juvenile.

 

Locality: Snyder Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

CM 1400: Maxilla, cranium and both lower jaws, two dorsal, seven caudal vertebrae, ribs, chevrons, part of pubis, ilium, femur and other bones.
Note: Maxilla in Fiorillo, 1991.

 

Locality: Lance Creek, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

CM 244: A phalanx.

GILMORE, 1920

USNM 2110: Metatarsal IV of the right foot missing proximal end.

 

Locality: Hager T-rex site, Garfield County, Montana

Horizon: Hell Creek.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Held at the Museum of the Rockies.

Material:

MOR 009 (Formerly GE-69-1): Articulated skull, eroded to the midline, medial surfaces of many of the elements are exposed, parts of both limbs, dorsal, caudal vertebrae, ilium, ischia, partial pubis, ribs, chevrons, partial feet and astragalus.

 

Locality: Location V75172, Shell Hill, Garfield County, Montana.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

UCMP 124488: Tooth.

 

Locality:?

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

MMS 51-2004: Left frontal.

DMNH 2827: Mounted skull and skeleton of a subadult.

 

Locality: Southeastern Montana.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

University of Wisconsin-Madison: 2/3 back part of the skull, pelvis, vertebrae, ribs and toe bones.

 

ANONYMOUS, 1993

Locality:

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

UCMP: Teeth marks on a piece of the pelvis of a Triceratops.

 

ANONYMOUS, 1994/TOKARYK, 1985, 2001, 2002, TOKARYK & BYRANT, 2004

Locality: SMNH Locality 72F07-0022, in the Frenchman River valley, 215 miles southwest of Regina, near the North Dakota Border, Saskatchewan Province, Canada.

Horizon: Frenchman Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.
Note: 28.3 m below the K-T boundary.

Material:

SMNH P2523.8: Fragmentary skull and skeleton.

Note: Nick named “Scotty.”

Note: A large coprolite was found near this specimen. Found in the coprolite are some bone fragments of a hadrosaur.

 

ANONYMOUS, 1997

DIS 101

Locality: South Dakota.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

DIS 101: Fragmentary skull, and fragmentary skeleton.

 

BAKKER, WILLIAMS & CURRIE, 1988

Locality: Corson County, South Dakota.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SMNH P2523.8: Five juvenile teeth.

 

BJORK, 1985

Locality: Ruby Site, Northeastern Butte County, South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

BRAMAN, 1988

Locality: Crows Nest, Alberta Province, Canada.

Horizon: Willow Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 81.6.1 (Black beauty): Skull and partial skeleton.

 

BURGE, 1997 (pers com)

Locality: Utah.

Horizon: North Horn Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth, fragmentary skull and fragmentary skeleton.

 

CARPENTER, 1979

Locality: Weld County, Colorado.

Horizon: Laramie Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCM 38804: Partial tooth.

 

= Aublysodon mirandus CARPENTER, 1982

Locality: UCMP V-5711, Bushy Tailed Blowout, Quarry area includes the floor of a small blowout approximately 10 yards northwest of the highest point on the ridge forming the northeastern side of the blowouts of loc. V-5003, S 1/2, Sec. 20, T37N, R64W, Lance Creek, Powder River Basin, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCM 43447: Tooth of a baby.

 

Locality: UCMP V5622, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCM 37878, 43447, UCMP 73101, 85141, 124993, 124995, 124996, 125229, 125230, 125233, 125234, 125237: Teeth.

UCMP 73091: Tooth of a baby.

 

Locality: UCMP V-5165, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 124367: Tooth of a baby.

 

Locality: UCMP V-5620, Lull Quarry, near the head of a tributary of Buck Creek, Powder River Basin, Niobrara County (Formerly Converse County), Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 124399: Tooth of a baby.

 

 

Locality: UCMP V-3087, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 124406, 124978, 124980, 124981, 124982: Teeth of babies.

UCMP 124979, 124994: Teeth.

 

 

Locality: Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

RTMP 87.112.33, 87.114.7: Teeth.

 

Locality: Arapahoe County, Colorado.

Horizon: Denver Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCM 38060: Tooth.

 

Locality: San Juan County, New Mexico.

Horizon: Naashoibito Member, Latest Kirtland Formation.

Biostratigraphy:

Age: Lancian age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

KU 12419: Tooth.

CARPENTER pers comm

Locality: Haystack Butte about 15 miles east of Hoover, Butte County, South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 12047: Skull and partial Skeleton.

 

 

Locality: Ruby Deposit, Northeastern Butte County, South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Field No:? SDSM 8354/PRB8775: Skull and much of the skeleton.

 

 

CARR & WILLIAMSON, 2000

Locality: New Mexico.

Horizon: Naashoibito Member, Latest Kirtland Formation.

Biostratigraphy:

Age: Lancian age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

NMMNH P-7199: Partial left dentary, tooth fragments and a partial vertebra.

 

CURRIE pers comm.

Locality: Dry Island Buffalo Jump Provincial Park, 40 miles north of the Tyrrell Museum of Paleontology, near Huxley, Alberta, Canada.

Horizon: Scollard Formation, Uppermost Edmonton Group.

Biostratigraphy:

Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

Never collected: Eroded skull.

RTMP 81.12.1: Skull and skeleton. (Huxley rex)

 

DERSTLER, 1994

Locality: Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Commercial collector: dorsal vertebrae.

1991 UNO Lance Dinosaur Expedition: A distal end of a juvenile metatarsal.

Private collector: Several vary large foot bones.

 

DERSTLER, 1995

Locality: Dragons’ Grave, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Shed teeth.

 

EIN, 1993

Locality: Remote ranch in Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Tip of a tooth found with the bones of “Kevin” Triceratops skeleton.

 

Locality: Dragon’s Grave, Remote ranch in Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Fragmentary teeth found with several Edmontosaurus skeletons.

ESTES, 1964

Locality: Powder River Basin, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Isolated fragments of teeth.

 

HECHT, 1997

Locality: Near Fort Peck, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Fragmentary skull and fragmentary skeleton.

 

HOLRYOD & HUTCHISON, 2002

Locality: V 86097, western North Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

172228:

 

Locality: V 86101, western North Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian Age, upper Maastrichtian Stage, uppermost Senonian Subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

172366:

 

HORNER pers comm.

Locality: Badlands, Montana.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

MOR: Fragmentary skull and fragmentary skeleton.
Note: This is a gracile animal.

 

JACOBSON & STROKA, 1995

Locality: Near Lemmon, Northwestern South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

University of Illinois: Shed tooth.

 

JANKE, 1996 & (SOTHERBY’S) ANONYMOUS, 1997

Locality: South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BOWAN: 10 percent of skeleton.

Z-rex (private collection): 50 percent of skeleton?

BHI-4100: 30 percent plus skeleton.

County rex: Partial skeleton.

STEVEN-9-14-95: Skeleton, still being excavated. Some eggshells found with skeleton.

South Dakota School of Mines & Technology Museum, Rapid City, South Dakota. 25% complete specimen.

 

Locality: Montana.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

University of Wisconsin: 10 percent of skeleton.

 

 

Locality: North Dakota.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Material:

Pioneer Trials Museum, Bowman, North Dakota: 5-10% complete.

 

LARSON & FREY, 1992

Locality: South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Black Hills Institute of Geological Research (Stan) 3033: Skull and skeleton.
Note: This is a gracile animal (Male).
Note: Broken ribs, ankylosed cervicals, bitten and broken parietal, anomoalous oppeings in the right jugal and surangular (MOLNAR, 2001).

 

LARSON, 1994

Locality: Harding County, South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

(Duffy): Some skull bones and fragmentary skeleton.

Not far from Stan and Duffy: A fragmentary skeleton.

 

LARSON, 1997 Pers. Com.

Locality: South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Kansas University:
Note: Male.

 

Locality: South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

BHI “Foxy”: Fragmentary skull and skeleton.
Note: Female nearly as large as “Sue”.

 

LANGSTON 1955 Field Notes.

Locality: Lac Pellebir-Eastend, Saskatchewan Province, Canada.

Horizon:

Biostratigraphy:

Age: Late Cretaceous.

Materiel:

Number: Not given: Toe Bone.

 

LANGSTON, 1955 Field Notes.

Locality: Frenchman river valley southeast of Eastend, Saskatchewan Province, Canada.

Horizon: Frenchman Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Toe Bones.

 

LAWSON, 1976/LEHMAN, 1985

Locality: Texas.

Horizon: Javelina Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

LERBEKMO, SINGH, JARZEN & RUSSELL, 1979/RUSSELL & SINGH, 1978

Locality: In a coulee tributary in the Red Deer River valley, 7 miles east of Huxley, NW 1/4, Sect. 10, Tp.34, R.22, W4, Alberta, Canada.

Horizon: 6m below a referred Triceratops albertensis and 10.5m below the Nevis coal seam, Scollard Formation, Uppermost Edmonton Group.

Biostratigraphy:

Age: Lancian Age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Upper Gulf Epoch, Late Cretaceous.

Material:

NMC 9554: Fragmentary cervical.

NMC 9950 (Now part of TMP 81.12.1): Pedal phalanx.

 

LOEWEN, SAMPSON, GETTY & DIFLEY, 2001, DIFLEY & EKDALE, 2002

Locality: Central Utah.

Horizon: North Horn Formation.

Biostratigraphy: Unit 1B

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UMNH 11000: Post orbital, and large tibia.

UMNH VP7513: Claw.

UMNH VP7626: Partial tooth.

 

LUCAS, MATEER, HUNT, & O'NEILL, 1987

Locality: San Juan Basin, New Mexico.

Horizon: Naashoibito Member, Latest Kirtland Formation.

Biostratigraphy:

Age: Lancian age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UNM FKK-076: Tooth.

 

 

LUPTON, GABRIEL & WEST, 1980

Locality: McCone County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Five teeth, a distal metapodial, a phalanx, and a vertebral fragment.

 

McMANNIS, 1965

Locality: Montana.

Horizon: Livingston Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

 

MOLNAR, 1978

Locality: ? Northwestern South Dakota.

Horizon: Lance Formation?

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MMS 51-2004 (Minnesota Museum of Science): Skull elements.

 

MOLNAR, 1991

Locality: Crooked Creek, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

AMNH 5005: Cranial fragments and fibula.
Note: Juvenile/subadult.

 

 

Locality: Unknown.

Horizon: Unknown.

Biostratigraphy:

Age:

Material:

AMNH 5881: Femur, tibia, fibula and phalanges.

 

Locality: T-rex skull site, Near Custer, Yellowstone county, Montana.

Horizon: Hell Creek Formation?

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MOR 008: Fragmentary skull.
Note: Palaeopathologies, pathological openings in the right surangulars, the squamosal articular surface of the right quadrate is rugose and partly hemispherical in form (MOLNAR, 2001).

 

Locality:

Horizon:

Biostratigraphy:

Age:

Material:

RTMP 82.50.11: Large right maxilla.

 

Locality: Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 131583: Maxilla and dentaries.

SCHMITT & SCHWEITZER, 2002

Locality: T-rex site, McCone County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

MOR 555: Skull and skeleton.
Note: Bloodcells?

 

SPENCER, TURNER & CHADWICK, 2001

Locality: Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Teeth.

 

STENERSON & O’CONNER, 1994

Locality: Northwestern South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given: Phalange of a juvenile.

 

TANKE pers comm.

Locality: Garfield County? Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UCMP 118742: Right maxilla of a very large Tyrannosaurus.

 

TYSON, 1997

Locality: Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

University of Notre Dame: Fragmentary skull and fragmentary skeleton.
Note: Pubis is 133 cm long, the article has Tyrannosaurus imperator?

 

WHITEMORE, 1988

Locality: SDSM Locality V851, Zerbst Locality, east of the Cheyenne River, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

SDSM 15115: 1 tooth.

SDSM 15117: 1 tooth fragment.

 

WOLBERG, LONZINSKY & HUNT, 1986

Locality: NMMNH Locality 353, Staton-LaPoint locality, 1.1 km north of Kettle Top Butte on the eastern shore of Elephant Butte Reservoir, New Mexico.

Horizon: Hall Lake Member, McRae Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

NMMNH P-3698 (old NMMNH P-10131-1): Left dentary, right palatine (misidentified as an incomplete articular), a right prearticular, incomplete teeth, and one nearly complete chevron.

 

WROBLEWSKI,1998

Locality: Western Hanna Basin, Carbon County, South-central Wyoming.

Horizon: Ferris Formation.

Biostratigraphy:

Age: Lancian age, uppermost Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

Number: Not given:

Also

= Aublysodon mirandus WROBLEWSKI,1998

 

= Manospondylus gigas COPE, 1892 (nomen dubium)
Etymology:

Holotype: AMNH: 3892
(Cope Collection)

Locality: Not given by Cope, But state verbally to Marsh and Lull that they were found in South Dakota.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Two dorsal centrum.

 

= Aublysodon amplus MARSH, 1892
Etymology:
= Deinodon amplus (MARSH 1892)
= Manospondylus amplus (MARSH 1892)
= Tyrannosaurus amplus (MARSH 1892)
= Stygivenator amplus (MARSH 1892) OLSHEVSKY, FORD & YAMAMOTO, 1995

Lectotype: YPM 296

Locality: Peterson's Quarry, Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Tooth.

 

= Aublysodon cristatus MARSH 1892
Etymology:
= Deinodon cristatus (MARSH 1892)
= Stygivenator cristatus (MARSH, 1892) OLSHEVSKY, FORD & YAMAMOTO, 1995

Lectotype: YPM 297

Locality: Niobrara County, Wyoming.

Horizon: Lance Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Tooth.

 

= Dynamosaurus imperiosus OSBORN, 1905
Etymology:
= Tyrannosaurus imperiosus OSBORN, 1906

Holotype: BMNH R7994 (formerly AMNH 5866)

Locality. Seven Mile Creek, six miles north of Cheyenne River, Weston County, Wyoming.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Right and left palatine, right and left dentary, fragmentary left femur, fragmentary right ilium, 2 sacral vertebrae and spines, atlas, axis, cervicals 3-9, cervico-dorsal 10-11, dorsal vertebrae.

 

= Genus nova LAWSON, 1972 (chironym)
Etymology:

Holotype: TMM 41436-1

Locality: Northeast part of the Big Bend National Park, Brewster County, Texas.

Horizon: Tornillo Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Left Maxilla.

 

= Albertosaurus cf lancensis MOLNAR, 1980
= Albertosaurus megagracilis PAUL, 1988
Etymology:
= Dinotyrannus megagracilis (PAUL, 1988) OLSHEVSKY, FORD & YAMAMOTO, 1995

Holotype: LACM 23845

Locality: On the Lester D. Engdahl Ranch, S34, T21N, R37E, near Jordan, Garfield County, Montana.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Nasals, fragmentary maxillae, fragmentary lachrymal, nearly complete frontals, braincase, anteromedial portion of the parietal, the medial portion of supraoccipital, fragmentary pterygoids, nearly complete quadratojugal, most of the right, but only fragments of the left surangular, left ulna lacking the radial and olecranal edges of the proximal articular surface, metacarpal II, manual ungual phalanx, proximal two-thirds of the left femur, proximal half of left tibia, right fibula, right astragalus, metatarsal II, distal half of metatarsal III, and pedal phalanges, (11 phalanges of the right pes are preserved, more or less complete.

 

 

= Tyrannosaurus stanwinstonorum PICKERING in prep
Note: Also SDSMT 12047

FMNH PR2081 (former Black Hills Institute of Geological Research “Sue” 2033)
= Tyrannosaurus stanwinstonorum PICKERING in prep

Locality: Maurice Williams’s Ranch, Cheyenne River Indian Reservation, near Faith, Hill City, South Dakota.

Horizon: Hell Creek Formation.

Biostratigraphy:

Age: Lancian age, upper Maastrichtian Stage, uppermost Senonian subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Skull and skeleton. With pathologies, 2 broken fibula, broken ribs with a tyrannosaur tooth in one, 2 broken caudal vertebrae, injured lower jaw with malformed double cusped tooth, and the top of the skull with tooth marks across it.
Note: Possible half a furcula BROCHU, 2003
Note: Pathologies are healed osteomylitis (infection) of the left fibula, healded proximal humerus, exuberant vertebral osteophyte formation, and multiple bilaterally-present erosive lesions primarily confined to the post-dentary mandibular elements. No evidence of pre or peri-mortem bite marks (REGA, 2001, BROCHU, (2002) 2003)

Found with Sue, a smaller adult male, a juvenile and infant.

A proximal tibia.

Partial fibula.

Partial tibia.

Cranial and front bones.

Note: This is a robust animal. (Female)