VETEROVATA
(Parataxonomy of fossil eggs and shells)

oofamily: Indet.

Oogenus: Oolithus (Oolithes) non BUCKMAN, 1859
O. sphaericus van STRAELEN, 1928
O. megadermus YOUNG, 1959
O. nanhsiungensis YOUNG, 1965

Gen. sp indet.

OOCLASS: AMPHIBIA LINNAEUS, 1758

OOSUBCLASS: LABYRINTHODONTIA OWEN, 1860

Incertae sedis

Oogenus: Mazonova GODFREY, 1995
M. helmichnus GODFREY, 1995 (Ootype)

Gen. sp indet.

OORDER: TESTUDINES NOVA
(Eggs with testudoid basic shell type)

Family: TESTUDOOLITHIDAE Hirsch, 1996

Oogenus: Oolithes BUCKMAN, 1859
O. bathonicoe BUCKMAN, 1859 (Ootype)
= Testudoflexoolithus bathonicae HIRSCH, 1996

Genus: Nova? O’BRIEN, 1995

Gen. sp indet.

Soft-shelled eggs of lizards and snakes
(Squamata)

Gen. sp indet.

Eggs with gekoid basic shell type
(Gekkota)

Gen. sp indet.

OORDER: CROCODYLINES NOVA

oofamily: KROKOLITHIDAE Kohring & Hirsch, 1996

Oogenus: Krokolithes HIRSCH, 1985
K. wilsoni HIRSCH, 1985 (Ootype)
K. helleri KOHRING & HIRSCH, 1996

Gen. sp indet.

OOSUPERCOHORT: DINOSAURIOVUM VIALOV, 1975
= GRUMULIOVUM VIALOV, 1986
= ORNATIOVUM VIALOV, 1986
Incertae sedis

Gen. Sp indet.

oofamily: DINOSAURIOVIDAE Vialov, 1986


Oogenus: Dinosauriovum VIALOV, 1975
Oosubgenus: Disparovum VIALOV, 1986
D. D. compositium VIALOV, 1986 (Ootype)

Oosubgenus: Grumuliovum VIALOV, 1986
D. G. tuberculatum VIALOV, 1986 (Ootype)
D. G. asperum VIALOV, 1986
D. G. punctatum VIALOV, 1986

Oosubgenus: Laeviovum VIALOV, 1986
D. L. laevis VIALOV, 1986 (Ootype)

Oosubgenus: Ornatiovum VIALOV, 1986
D. O. multifarium VIALOV, 1986 (Ootype)
D. O. partentosum VIALOV, 1986
D. O. subtilis VIALOV, 1986

OOCOHORT: THEROPODA NOVA

EGGSHELLS OF THE ORNITHOID BASIC TYPE THEROPODA

Incertae sedis

Gen. sp indet.

Family: COMPSOGNATHIDAE Cope, 1875

Genus: Compsognathus WAGNER, 1859
C. longipes WAGNER, 1859

Genus: Sinosauropteryx JI QIANG et. al., 1996
S. prima JI QIANG, et al., 1996

Family: THERIZINOSAURIDAE Maleev, 1954

Genus: Nova? MANNING, JOYSEY, & CRUICKSHANK, 1997, 2000

oofamily: ELONGATOOLITHIDAE Zhao, 1975

Note: Elongatoolithids are known to belong to theropod dinosaurs based on new material collected in Mongolia and China. The well-preserved remnants of an oviraptorid embryo have been found within an Elongatoolithus egg (Norell et al, 1994). Oviraptorosaur embryos have also been found in Macroelongatoolithus eggs from Henan Province, China (Zelenitsky, pers. comm.). Additionally, adult specimens of Oviraptor have been found atop of nests of Elongatoolithus eggs (Dong and Currie, 1996; Norell et al., 1995). Elongatoolithid eggshells are also known from Lower Cretaceous strata of North America (Zelenitsky pers. comm.); These rocks also contain the remains of probable Oviraptorosaurs (Makovicky and Sues, 1998).

Oogenus: Ellipsoolithus MOHABEY, 1998
E. khedaensis MOHABEY, 1998 (Ootype)

Oogenus: Elongatoolithus ZHAO, 1975
E. elongatus (YOUNG, 1954) ZHAO, 1975 (Ootype)
= Oolites elongatus YOUNG, 1954
E. andrewsi ZHAO, 1975
E. magnus ZENG & ZHANG, 1979
E. frustrabilis MIKHAILOV, 1994
E. subtitectorius MIKHAILOV, 1994
E. sigillarius MIKHAILOV, 1994
E. excellens MIKHAILOV, 1994
E. sp

Elongatolithus?

Genus: Oviraptor OSBORN, 1924
O. philoceratops OSBORN, 1924

Genus: Ingenia BARSBOLD, 1981
I. sp

Oogenus: Heishanoolithus ZHAO & ZHAO, 1999
H. changii ZHAO & ZHAO, 1999 (Ootype)

Oogenus: Lepidotoolithus ZHANG & XUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996)
L. guofenglouensis ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996) (Ootype)

Oogenus: Macroelongatoolithus LI, AN, ZHU, ZHANG, LIU, QU, YOU, LIANG, LI, et al.,1995
M. carlylei (JENSEN, 1970) ZELENITSKY, CARPENTER & CURRIE, 2000
= Oolithes carlyensis JENSEN, 1970
= Oolithes carliensis MIKHAILOV, 1991 (sic)
= Spheroolithus carlyensis (JENSEN, 1970)
= Boletuoolithis carlyensis (JENSEN, 1970) ZELENITSKY, CARPENTER & CURRIE, 2000
M. xixiaensis LI, AN, ZHU, ZHANG, LIU, QU, YOU, LIANG, LI, et al.,1995 (Ootype)
= Longiteresoolithus xixiaensis WANG & ZHOU, 1995

Oogenus: Macroolithus ZHAO, 1975
M. rugustus (YOUNG, 1965) ZHAO, 1975 (Ootype)
= Oolithes rugustus YOUNG, 1965
M. yaotunensis ZHAO, 1975
M. mutabilis MIKHAILOV, 1994
M. turolensis AMO SANJUAN, CANUDO & CUENCA BESCOS, 2000
M. sp

Oogenus: Nanhsiungoolithus ZHAO, 1975
N. chuetienensis ZHAO, 1975 (Ootype)

Oogenus: Spongioolithus BRAY, 1999
S. hirschi BRAY, 1999 (Ootype)

Oogenus: Trachoolithus MIKHAILOV, 1994
T. faticanus MIKHAILOV, 1994 (Ootype)

Gen. Sp indet.

Family: TYRANNOSAURIDAE Osborn, 1906

Subfamily: TYRANNOSAURINAE Osborn, 1906

Tribe: TYRANNOSAURINI Osborn, 1906 (sensu Olshevsky, Ford & Yamamoto, 1995)

Tyrannosaurus rex

Family: DROMAEOSAURIDAE Russell, 1969

Subfamily: VELOCIRAPATORINAE Barsbold, 1978

Eggshells associated with Deinonychus antirrhopus (MAKOVICKY & GRELLET-TINNER, 2000)

Family: TROODONTIDAE Gilmore, 1924

Troodon formosus

oofamily: OBLONGOOLITHIDAE Mikhailov, 1996

Oogenus: Oblongoolithus MIKHAILOV, 1996
O. glaber MIKHAILOV, 1996 (Ootype)

EGGS WITH DINOSAUROID-PRISMATIC BASIC SHELL TYPE

oofamily: PRISMATOOLITHIDAE Hirsch, 1994

Note: Prismatoolithid eggs from the Upper Cretaceous of Montana were initially attributed to hypsilophodontid dinosaurs, based on embryonic remains within eggs (Horner and Weishampel, 1888; Hirsch and Quinn, 1990). Subsequently, numerous prismatoolithid egg fossils from the Santonian-Campanian of Mongolia (oogenus Protoceratopsidovum) were attributed to protoceratopsian dinosaurs, based on the common occurrence of protoceratopsids in these strata (Mikhailov, 1994a; Mikhailov et al, 1994). It has been announced recently that assignment of the embryonic remains from the prismatoolithid eggs from Montana to the hypsilophodontid Orodromeus makelai was erroneous. The embryo has since been reassigned to the theropod Troodon cf. formosus (Horner and Weishampel 1996), Prismatoolithid eggs share several characteristics with other theropod eggs (e.g., elongatoolithids) and their initial assignment to hypsilophodonts and protoceratopsids is probably incorrect (Zelenitsky, pers. comm.).

Oogenus: Oolithus (Oolithes) non BUCKMAN, 1859
O. laminadermus CHAO & CHANG, 1974

Oogenus: Preprismatoolithus ZELENITSKY & HILLS, 1996
= Prismatoolithus HIRSCH, 1994
P. coloradensis (HIRSCH, 1994) ZELENITSKY & HILLS, 1996 (Ootype)
= Prismatoolithus coloradensis HIRSCH, 1994

Oogenus: Prismatoolithus ZHAO & LI, 1993
P. gebiensis ZHAO & LI, 1993 (Ootype)
P. tenuis VIANEY-LIAUD & CROCHET, 1993
P. matellensis VIANEY-LIAUD & CROCHET, 1993
P. levis ZELENITSKY & HILLS, 1996
P. jenseni BRAY, 1999
P. hukouensis ZHAO, 1999
P. caboti GGARCIA, FEIST, CABOT, VALENTIN & VIANEY-LIAUD, 2000
P. Oospecies: Nova VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
P. sp

Oogenus: Protoceratopsidovum MIKHAILOV, 1994
P. sincerum MIKHAILOV, 1994 (Ootype)
P. minimum MIKHAILOV, 1994
P. fluxuosum MIKHAILOV, 1994

Oogenus: Spheruprismatoolithus BRAY, 1999
S. condensus BRAY, 1999 (Ootype)

Gen. sp indet.

MORPHOTYPE: ORNITHOID-PRISMATIC INCERTAE SEDIS

Oogenus: Dispersituberoolithus ZELENITSKY, HILLS & CURRIE, 1996
D. exilis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)

Oogenus: Pseudogeckoolithus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
P. nodosus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997 (Ootype)
Note: Morphotype: Geckonoid?
P. sp

OOCOHORT: ORNITHISCHIFORMES NOVA

OOSUPERORDER: SAUROPODOMORPHA NOVA

TUBOSPHERULITIC STRUCTURAL MORPHOTYPE

oofamily: "MEGALOOLITHIDAE" Zhao, 1979
[Sauropoda]

Note: The first dinosaur eggs remains were discovered in Cretaceous strata of Southern France over a century ago (Pouech..). These eggs, known as megaloolithids, have been assumed to belong to sauropod dinosaurs because they are usually found in strata that are rich in sauropod remains (Erben et al., 1979; Vianey-Liaud et al. 1987; Jain 1989; Sahni et al, 1994). Recent confirmation of this correlation was made with the discovery of South American megaloolithid eggs containing sauropod embryos (Chiappe et al, 1998). Occasional references to typical megaloolithid eggshells as “hadrosaurian’ (Grigorescu et al, 1994, p. 77) are obviously misleading. Additionally, Upper Cretaceous strata in North America are rich hadrosaur remains and contain no trace of megaloolithid eggshell (Zelenitsky, pers. comm.). All positively identified hadrosaurian eggs have a different eggshell structure (see Spheroolithidae).

Oogenus: Cairanoolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
= Dughioolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
C. dughii VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994 (Ootype)
C. roussetensis (VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994) BARCIA & VIANEY-LIAUD, 2001
= Dughioolithus roussetensis VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994

Oogenus: Megaloolithus VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. mammilare VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994 (Ootype)
M. siruguei VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. aurelinesis VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. petralta VIANEY-LIAUD, MALLAN, BUSCALI & MONTGELARD, 1994
M. trempii VIANEY-LIAUD, MALLAN, BUSCAIL & MONTGELARD, 1994
M. cylindricus KHOLSA & SAHNI, 1995
= Megaloolithus rahioliensis MOHABEY, 1998
M. jabalpurensis KHOLSA & SAHNI, 1995
= Megaloolithus patagonicus CALVO, ENGLLAND, HEREDIA & SALGADO, 1997
= Megaloolithus matleyi MOHABEY, 1997
M. mohabeyi KHOLSA & SAHNI, 1995
= Megaloolithus phensaniensis MOHABEY, 1998
M. baghensis KHOLSA & SAHNI, 1995
= Megaloolithus pseudomamillare VIANEY-LIAUD, HERISCH, SAHNI & SIGE, 1997
= Megaloolithus balasinorensis MOHABEY, 1998
M. dholiyaensis KHOLSA & SAHNI, 1995
M. padiyalensis KHOLSA & SAHNI, 1995
M. walpurensis KHOLSA & SAHNI, 1995
= Megaloolithus khempurensis MOHABEY, 1998
Note: VIANEY-LIAUD, KHOSLA & GARCIA, 2003 place Megaloolithus walpurensis within the M. khempurensis because of the fragmentary nature of M. walpurensis, but M. walpurensis has priority.
M. dhoridungriensis MOHABEY, 1998
M. megadermus MOHABEY, 1998
M. microtuberculata GARCIA & VIANEY-LIAUD, 2001
M. sp

Gen. sp indet.

Incertae sedis

Oogenus: Tacuaremborum MONES, 1980
T. oblongum MONES, 1980 (Ootype)

oofamily: FAVEOLOOLITHIDAE Zhao & Ding, 1976 [Sauropoda]

Note: MIKHAILOV, 1997…Only bones of sauropod dinosaurs are known from the Faveoloolithus nest-bearing layers in Ologoy-Ulan-Tsav locality in Mongolia Gobi (Sochava, 1969; S. M. Kurzanov, pers. comm.). The arrangement of the eggs and porosity of the eggshell appear to be similar to those subterranean nesters, such as chelonian.

Oogenus: Faveoloolithus ZHAO & DING, 1976
F. ningxiaensis ZHAO & DING, 1976 (Ootype)
F. sp

Oogenus: Sphaerovum MONES, 1980
S. erbeni MONES, 1980 (Ootype)

Oogenus: Youngoolithus ZHAO 1979
Y. xiaguanensis ZHAO, 1979(Ootype)
Y. xipingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998

Gen. sp indet.

Family: Unknown.

Gen. sp indet.

Family: Titanosauridae?

Hypselosaurus priscus?

Gen. sp indet.

oofamily: DENDROOLITHIDAE Zhao and Li, 1988
[Sauropoda? Therizinosauria? Mikhailov, 1997

Note: As per Mikhailov, 1997…Based on the taphonomic settings and reconstructed type of nesting, dendroolithid remains have been attributed tentatively to some ornithopod or sauropod dinosaurs (Mikhailov, 1991). Recently, some privately owned dendroolithid eggs from Henan Province, China displayed at the University Museum of Zoology, Cambridge (the author was kindly allowed to examine shell samples) have been shown to contain well preserved embryos of therizinosaurid dinosaurs at late stages of development (Cohen et al, 1995)…

Oogenus: Dendroolithus ZHAO & LI, 1988
D. wangdianensis ZHAO & LI, 1988 (Ootype)
D. verrucarius MIKHAILOV, 1994
D. microporosus MIKHAILOV, 1994
D. xichuanensis ZHAO & ZHAO, 1998
D. furcatus XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. dendriticus XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. sanlimiaoensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
D. zhaoyingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998

Oogenus: Placcolithus ZHAO & ZHAO, 1998
P. taohensis ZHAO & ZHAO, 1998 (Ootype)

Gen. sp indet.

oofamily: DICTYOOLITHIDAE Zhao, 1994

Oogenus: Dictyoolithus ZHAO, 1994
D. hongpoensis ZHAO, 1994 (Ootype)
D. neixiangensis ZHAO, 1994
D. jiangi LIU& ZHAO, 2004

Gen. sp indet.

oofamily: PHACELOOLITHIDAE Zeng& Zhang, 1979

Oogenus: Phaceloolithus ZENG & ZHANG, 1979
P. huanensis ZENG & ZHANG, 1979 (Ootype)

oofamily: Incertae sedis ?spherulite or spheruprismatic structural type eggshell.

Oogenus: Parvoolithus MIKHAILOV, 1996
P. tortuosus MIKHAILOV, 1996 (Ootype)
Note: Possible beetle castings ?

Gen. sp indet.

oofamily: Incertae sedis

Oogenus: Placoolithus non ZHAO, 1979
P. cf. taohensis ZHAO, 1979 (Ootype)

Oogenus: Stromatoolithus ZHAO, YE, LI, ZHAO & YAN, 1991
S. pinglingensis ZHAO, YE, LI, ZHAO & YAN, 1991 (Ootype)

OOMAGNORDER: ORNITHISCHIA SEELEY, 1888
EGGS WITH DINOSAUROID-SPHERULITIC BASIC SHELL TYPE

oofamily: SPHEROOLITHIDAE Zhao, 1979
[Ornithopoda: some hadrosaurs]

Note: As per Mikhailov, 1997…Spheroolithus eggs from Laiyang have been attributed to hadrosaurs (Young, 1959, pp. 511, 517); the eggs of S. irenensis were found in close association with Bactrosaurus remains (Zhao and Jiang, 1974, p. 81). Spheroolithid eggs from the Upper Cretaceous Two Medicine Formation of Montana were found in association with hadrosaurian juveniles (Horner and Makela, 1979). Lambeosaurine embryos (Hypacrosaurus stebingeri; Horner and Currie, 1994) were also found in association with large spherical eggs; however, the eggshell morphotype could not be identified as spheroolithid because it is diagenetically modified (Zelenitsky, 1995). This was misinterpreted by Mikhailov. These discoveries have allowed all spheroolithid remains to be attributed to hadrosaurs (Hirsch and Quinn, 1990, Mikhailov et al, 1994). Their abundance, as well as their geographical and stratigraphical distribution, are in accordance with this conclusion…

Oogenus: Paraspheroolithus ZHAO, 1979
Note: (= Spheroolithus ZHAO, 1979 as per MIKHAILOV, 1997)
P. irenensis (CHAO & CHIANG, 1974) ZHAO, 1979 (Ootype)
= Spheroolithus irenensis (CHAO & CHIANG, 1974) MIKHAILOV, 1994
= Oolithes spheroides YOUNG, 1954
= Oolithes irenensis CHAO & CHIANG, 1974
= Spheroolithus sp MIKHAILOV, 1991
P. lamelliformae ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996)
P. shizuiwanensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
P. yangchengensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
P. sp

Oogenus: Spheroolithus ZHAO, 1979
S. spheroides (YOUNG, 1954) ZHAO, 1979 (Ootype)
= Oolithes spheroides YOUNG, 1954
S. chiangchiungtingensis (CHAO & CHIANG, 1974) ZHAO, 1979
= Oolithes chiangchiungtigensis CHAO & CHIANG, 1974
S. megadermus (YOUNG, 1959) ZHAO, 1979
= Oolithes megadermus YOUNG, 1959
S. tenuicorticus MIKHAILOV, 1994
S. maiasauroides MIKHAILOV, 1994
S. albertensis ZELENITSKY & HILLS, 1997
Oosp (unnamed)
= Genus: Maiasaura HORNER & MAKELA, 1979
M. peeblesorum HORNER & MAKELA, 1979
S. sp

Oogenus: Shangyangoolithus ZHANG & XUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996)
= Shanyangoolithus ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996) (sic)
S. shanyangensis ZHANG & ZUE, 1999 (in XUE, ZHANG, BI, YUE & CHEN, 1996) (Ootype)

Oogenus: Shixingoolithus ZHAO, YE, LI, ZHAO, & ZHANG, 1991
S. erbeni ZHAO, YE, LI, ZHAO, & ZHANG, 1991 (Ootype)

Genus: Nova XU, ZHAO, LU, DONG & HUANG, 1999

Gen. sp indet.

oofamily: OVALOOLITHIDAE Mikhailov, 1991
(?Ornithopoda)

Note: As per Mikhailov, 1997…The histrostructure of the shell and other, superficial features of the egg, are similar to those of the Spheroolithidae. At the Mogoyn-Ulagiyn-Kheates locality (= Tel-Ulan-Ula; Mongolia, Dornogov Aimak), pinkish strata containing numerous ovaloolithid eggs in clutches yield remnants of only lambeosaurine dinosaurs (S. M. Kurzanov, pers. comm.). This prompts the tentative assignment of ovaloolithid remains to ornithopod dinosaurs. Preliminary assignment of embryonic limb bones associated with an ovaloolithid egg (PIN 2970/9) to protoceratopsian dinosaurs (L. I. Hozatcky in Sochen, 1972) has proved to be incorrect (S. M. Kurzanov, see Mikhailov et al., 1994)…

Oogenus: Ovaloolithus ZHAO, 1978
O. chinkangkouensis (CHAO & CHIANG, 1974) ZHAO, 1978 (Ootype)
= Oolithes chinkangkouensis CHAO & CHIANG, 1974
O. laminadermus (CHAO & CHIANG, 1974) ZHAO, 1979
= Oolithes sp YOUNG, 1954
= Oolithes laminadermus (YOUNG, 1954) CHAO & CHIANG, 1974
O. tristriatus ZHAO, 1979
O. mixtistriatus ZHAO, 1979
O. monostriatus ZHAO, 1979
O. dinornithoides MIKHAILOV, 1994
O. sangpingensis XIAOSI, LIWU, CHENG, ZOU, PANG, QANG, CHEN, ZHEN, WANG, LIU, XIE & JIN, 1998
O. utahensis BRAY, 1999
O. tenuisus BRAY, 1999
O. sp

Gen. sp indet.

Ornithischia incertae sedis?

HYPSILOPHODONTIDAE

Gen. sp indet.

RHABDODONTIDAE

Gen. sp indet.

DRYOSAURIDAE

Gen. sp indet.

CAMPTOSAURIDAE

Gen. sp indet.

HADROSAURIDAE

Telmatosaurus transsylvanicus

Gen. sp indet.

LAMBEOSAURINAE

Hypacrosaurus stebingeri

PSITTACOSAURIDAE

Gen. sp indet.

PROTOCERATOPIDAE?

Protoceratops

Gen. sp indet.

KENTROSAURINAE

Dacenterurus armatus

OOCLASS: AVES NOVA

Incertae sedis

Gen. Sp indet.

oofamily: LAEVISOOLITHIDAE Mikhailov, 1991 [AVES, Enantiornithes]

Note: As per Mikhailov, 1997… These small and thin-shelled eggs with ornithoid eggshell structure were proposed to belong either to birds or small theropod dinosaurs (Mikhailov, 1991). Recently, one of the specimens of laevisoolithid eggshells was found in situ with an associated partial skeleton of an enantiornithid bird Nanantius (Kurockhin, 1996). This allows taxonomic assignment of laevisoolithid eggs to Enantiornithes (Mikhailov, 1996a)…

Oogenus: Laevisoolithus MIKHAILOV, 1991
L. sochavai MIKHAILOV, 1991 (Ootype)

Genus: Gobipteryx ELZANOWSKI, 1976
G. minuta ELZANOWSKI, 1976
= Nanantius valifanovi KUROCHKIN, 1996
G. sp

Gen. sp indet.

oofamily: GOBIOOLITHIDAE Mikhailov, 1996 [Aves]

Note: As per Mikhailov, 1997…Well-developed embryos of palaeognathous birds have been found at Khermiyn-Tsav (Elzanowski, 1981; Kurochkin, 1988). Although Elzanowski never stated that these embryos should be assigned to the small bird Gobipteryx minuta, described earlier from deposits of the same age (Elzanowski, 1977), this opinion has occasionally appeared in the literature Recent re-examination of embryonic material (Kurockhin, 1995) has shown that the embryo, which shares some features with Gobipteyx, is not certainly associated with the Gobioolithus minor eggs. Embryos that are certainly from Gobioolithus minor eggs may belong to a different group of birds, apparently Palaeognathae…

Genus: Gobipipus CHATTERJEE, 1994 (noma nudum)
G. elzanowskii CHATTERJEE, 1994 (noma nudum)

Oogenus: Gobioolithus MIKHAILOV, 1996
G. minor MIKHAILOV, 1996 (Ootype)
G. major MIKHAILOV, 1996

oofamily: SUBTILIOLITHIDAE Mikhailov, 1991 [AVES]

Oogenus: Subtiliolithus MIKHAILOV, 1991
S. microtuberculatus MIKHAILOV, 1991 (Ootype)
S. kachchhensis KHOLSA & SAHNI, 1995

Morphotype: Ornithoid-ratite

Incertae sedis.

Oogenus: Ageroolithus VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997
A. fontllongensis VIANEY-LIAUD & LOPEZ-MARTINEZ, 1997 (Ootype)
A. sp

Oogenus: Continuoolithus ZELENITSKY, HILLS & CURRIE, 1996
C. canadensis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably small theropod.

Oogenus: Porituberoolithus ZELENITSKY, HILLS & CURRIE, 1996
P. warnerensis ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably small theropod.

Oogenus: Tristraguloolithus ZELENITSKY, HILLS & CURRIE, 1996
T. cracioides ZELENITSKY, HILLS & CURRIE, 1996 (Ootype)
Note: Probably belongs to a bird.

Gen. sp indet.

PTEROSAURIA?

Gen. sp indet.

Bibliography

Bibliography for taxa list

A

Amo Sanjuan O., Canudo J. I., and Cuenca Bescos G., 2000, First record of elongatoolithid eggshells from the Lower Barremian (Lower Cretaceous) of Europe (Cuesta Corrales 2, Galve Basin, Teruel, Spain): In: First International Symposium on dinosaur eggs and babies, extended abstracts, edited by Bravo A. M., and Reyes T., 2000, p. 7-14.

B

Bray, E. S., 1998, Dinosaur Eggshell Boletuoolithus carlylensis, Oogenus nov. from the Lower Cretaceous Cedar Mountain Formation of Utah: In: Lower and Middle Cretaceous Terrestrial Ecosystems. Edited by Lucas S. G., Kirkland J. I., and Estep J. W., New Mexico Museum of Natural History and Science, Bulletin 14, p. 221-224.

Bray, E. S., 1999, Eggs and eggshell from the Upper Cretaceous North Horn Formation, Central Utah:
In: Vertebrate Paleontology in Utah. Edited by Gillette D. D., Miscellaneous Publication 99-1, Utah Geological Survey, 360-375.
Buckman, J., 1859, On some fossil reptilian eggs from the Great Oolite of Cirenchester: Proceedings of the Geological Society of London, 1859, p. 107-110.

C

Calvo, J. O., Englland S., Heredia S. E., and Salgado L., 1997, First record of dinosaur eggshells (?Sauropoda – Megaloolithidae) from Neuquen, Patagonia, Argentina: Gaia, n. 14, p. 23-32.

Chiappe, L. M., Coria R. A., Dingus L., Jackson F., Chinsamy A., and Fox M., 1998, Sauropod dinosaur embryos from the Late Cretaceous of Patagonia: Nature, v. 396, p. 258-261.

Chao, T.–K., and Chiang Y.-K., 1974, Microscopic studies on the dinosaurian egg-shells from Laiyang, Shantung Province: Scientia Sincia, v. 17, p. 71-83.

D

Dong, Z.-M., and Currie P. J., 1995, On the discovery of an Oviraptorid skeleton on a nest of eggs: Fifty-Fifth Annual Meeting Society of Vertebrate Paleontology, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, November 1-4, 1995 Journal of Vertebrate Paleontology, v. 15, Supplement to n. 3, Abstracts of Papers, p. 26a.

Dong, Z.-M., and Currie P. J., 1996, On the discovery of an oviraptorid skeleton on a nest of eggs at Bayan Mandahu, Inner Mongolia, People’s Republic of China Canadian Journal of Earth Science, v. 33, p. 631-636.

E

Erben, H. K., Ashraf A. R., Krumsiek K., and Thein J., 1983, Some Dinosaurs survived the Cretaceous “Final Event”: Terra Cognita, v. 3, n. 2, p. 211-212.

G

Garcia, G., Feist M., Cabot A., Valentin X., and Vianey-Liaud M., 2000, Les œufs de dinosaures du Cretace superieur du bassin de Villeveyrac-Meze (Herault, France): description d’une nouvelle ooespece de Prismatollithus: Bulletin de la societie geologiques de France, v. 171, n. 3, p. 283-289.

Garcia, G., and Vianey-Liaud, M., 2001, Nouvelles donnees sur les coquilles d’œufs de dinosaures Megaloolithidae du Sud de la France: systematiue et variabilite intraspecifique: Compte rendu hebdomadaire des seances de l’Academie des Sciences de la Terre et des plantes, v. 332, p. 183-191.
Godfrey, S. J., 1995, Fossilized eggs from the Pennsylvanian of Illinois: Ichnos, v. 4, p. 71-75.

H

Hirsch, K. F., 1985, Fossil Crocodilian eggs from the Eocene of Colorado: Journal of Paleontology, v. 59, n. 3, p. 531-534.

Hirsch, K. F., 1994, Upper Jurassic eggshells from the Western Interior of North America: In: Dinosaur Eggs and Babies, Edited by Carpenter K., Hirsch K. F., and Horner J. R., Cambridge University Press, p. 137-150.

Hirsch, K. F., 1996, Parataxonomic classification of fossil chelonian and gecko eggs: Journal of Vertebrate Paleontology, v. 16, n. 4, p. 752-762.

Hirsch K. F., and Quinn B., 1990, Eggs and eggshell fragments from the Late Cretaceous Two Medicine Formation of Montana: Journal of Vertebrate Paleontology, v, 10, n. 4, p. 491-511.

Horner, J. R., and Currie P. J., 1994, Embryonic and neonatal morphology and ontogeny of a new species of Hypacrosaurus (Ornithischia, Lambeosauridae) from Montana and Alberta: In: Dinosaur Eggs and Babies, Edited by Carpenter K., Hirsch K. F., and Horner J. R., Cambridge University Press, p. 312-336.

Horner, J. R., and Makela R., 1979, Nest of juveniles provides evidence of family structure among dinosaurs: Nature, v. 282, n. 5736, p. 296-298.

Horner, J. B., and Weishampel D. B., 1988, A comparative embryological study of two ornithischian dinosaurs: Nature, v. 332, p. 256-257.

Horner, J. B., and Weishampel D. B., 1996, A comparative embryological study of two ornithischian dinosaurs—a correction: Nature, v. 383, p. 103.

J

Jain, S. L., 1989, Recent Dinosaur Discoveries in India, Including Eggshells, Nests and Coprolites: In: Dinosaur Tracks and Traces, Edited by Gillette D. D., and Lockley M. G., Cambridge University Press, 99-108.

Jensen, J. A., 1970, Fossil Eggs in the Lower Cretaceous of Utah: Brigham Young University Geology Studies, v. 17, n. 1, p. 51-65.

K

Khosla, A., and Sahni A., 1995, Parataxonomic classification of Late Cretaceous Dinosaur Eggshells from India: Journal of The Palaeontological Society of India, v. 40, p. 87-102.

Kohring, R., and Hirsch K. F., 1996. Crocodilian and avian eggshells from the Middle Eocene of the Geiseltal, Eastern Germany: Journal of Vertebrate Paleontology, v. 16, n. 1, 67-80.

L

Li, Y., An C., Zhu, Y., Zhang Y., Liu Y., Qu L., You L., Liang X., Li X., et al, 1995, DNA isolation and sequence analysis of dinosaur DNA from Cretaceous Dinosaur egg in Xixia, Henan, China: Acta Scientiarum Naturalium Universitatis Pekinensis, v. 31, n. 2, p. 148-152.

Liu, J.-Y., and Zhao, X.-K., 2004, A new oospecies of the dinosaur eggs (Dictyoolithus) from Laiyang, Shandong Provine: Vertebrata PalAsiatica, v. 42, n. 2, p. 166-170.

Loyal, R. S., Mohabey D. M, Kholsa A., and Sahni A., (1998) 2000, Status and palaeobilogy of the Late Cretaceous Indian theropods with description of a new theropod eggshell oogenus and oospecies, Ellipsoolithus khedaensis, from the Lameta Formation, District Kheda, Gujarat, Western India: In: Aspects of Theropod Paleobiology, edited by Perez-Moreno B. P., Holtz T. Jr., Sanz J. L., and Moratalla J., Gaia, n. 15, p. 379-387.

M

Makovicky, P. J., and Sues H.-D., 1998, Anatomy and Phylogenetic Relationships of the Theropod Dinosaur Microvenator celer from the Lower Cretaceous of Montana: American Museum Novitates, n. 3240, p. 1-27.

Mikhailov, K. E., 1994, Theropod and Protoceratopsian Dinosaur eggs from the Cretaceous of Mongolia and Kazakhstan: Paleontological Journal, v. 28, n. 2, p. 101-120.

Mikhailov, K. E.. 1994, Eggs of Sauropod and Ornithopod Dinosaurs from the Cretaceous deposits of Mongolia: Paleontological Journal, v. 28, n. 3, p. 141-159.

Mikhailov, K. E., 1996, New genera of fossil eggs from the Late Cretaceous of Mongolia: Paleontological Journal, v. 30, n. 2, p. 246-248.

Mikhailov, K. E., 1996, Bird Eggs in the Late Cretaceous of Mongolia: Paleontological Journal, v. 30, n. 1, p. 114-116.

Mikhailov, K. E., 1997, Fossil and recent eggshell in amniotic vertebrates: Fine structure, comparative morphology and Classification: Special Papers in Paleontology, n. 56, p. 1-79.

Mohabey, D. M., 1996, A new oospecies, Megaloolithus matleyi, from the Lameta Formation (Late Cretaceous) of Chandrapur district, Maharashtra, India, and general remarks on the palaeoenvironment and nesting behavior of dinosaurs: Cretaceous Research, v. 17, p. 183-196.

Mohabey, D. M., 1998, Systematics of Indian Upper Cretaceous dinosaur and chelonian eggshells: Journal of Vertebrate Paleontology, v. 18, n. 2, p. 348-362.


Mones, A., 1980, Nuevos elementos de la paleoherpetofauna del Uruguay (Crocodilia y Dinosauria): Actas 2nd Congreso Argentino de Paleontologie y Bioestratigrafia y 1st Congreso Latinoamerica, (Buenos Aires), v. 1, p. 265-277.

N

Norrell, M. A., Clark J. M., Chiappe L. M., and Dashzeveg D., 1995, A nesting dinosaur: Nature, v. 378, p. 774-776.

S

Sahni, A., Tandon S. K., Jolly A., Bajpai S., Sood A., and Srinivasan S., 1994, Late Cretaceous dinosaur eggs and nesting sites from the Deccan valcano-sedimentary province of peninsular India. In: Dinosaur Eggs and Babies, Edited by Carpenter K., Hirsch K. F., and Horner J. R., Cambridge University Press, p. 204-226.Straelen, V. van, 1928, Les oeufs de reptiles fossils: Palaeontologica, v. 1, p. 295-317.

V

Vianey-Liaud, M., Hirsch K., Sahni A., and Sige B., 1997, Late Cretaceous Peruvian Eggshells and their relationships with Laurasian and Eastern Gondwanian material: Geobios, v. 30, n. 1, p. 75-90.

Vianey-Liaud, M., Jain S. L., and Sahni A., 1987, Dinosaur eggshells (Saurischia) from the Late Cretaceous Intertrappean and Lameta Formations (Deccan, India): Journal of Vertebrate Paleontology, v, 7, n. 4, p. 408-424.

Vianey-Liaud, M., and Lopez-Martinez N., 1997, Late Cretaceous Dinosaur Eggshells from Temp Basin, Southern Pyrenees, Lleida, Spain: Journal of Paleontology, v. 71, n. 6, p. 1157-1171.

Vianey-Liaud, M., Malln P., Buscali O., and Montgelard C., 1994, Review of French dinosaur eggshells; morphology, structure, mineral, and organic composition: In: Dinosaur Eggs and Babies, Edited by Carpenter K., Hirsch K. F., and Horner J. R., Cambridge University Press, p. 151-183.

Vianey-Liaud, M., Jain S. L., and Sahni A., 1987, Dinosaur eggshells (Saurischia) from the Late Cretaceous Intertrappean and Lameta Formations (Deccan, India): Journal of Vertebrate Paleontology, v. 7, n. 4, p.408-424.

X

Xiaosi, F., Lu L., Cheng Z., Zou Y., Pang O., Wang W., Chen K., Zhen Y., Wang X., Liu J., Xie H., and Jin Y., 1998, On the Cretaceous Fossil Eggs of Xixia County, Henan Province: Geological Publishing House Beijing, p. 1-125.

Xue, X., Zhang Y., Bi Y., Yue L., and Chen D., 1996.,The development and environmental changes of the Intermontane basins in the eastern part of Qinling Mountains: Geological Publishing House, Beijing, p. 1-181.

Y

Young, C.-C., 1954, Fossil Reptilian eggs from Laiyang, Shantung, China: Scientia Sincia, v. 3, n. 4, p. 505-522.

Young, C.-C., 1959, On a new fossil egg from Laiyang, Shantung: Vertebrata PalAsiatica, v. 3, n. 1, p. 34-35.

Young, C.-C., 1965, Fossil eggs from Nanhsiung, Kwangtung and Kanchou, Kiangsi: Vertebrata PalAsiatica, v. 9, n. 2, p. 141-170.

Z

Zeng, D., and Zhang J., 1979, On the dinosaurian eggs from the westen Dongting Basin, Hunan: Vertebrata PalAsiatica, v. 17, p. 131-136.

Zelenitsky, D. K., Carpenter K., and Currie P. J., 2000, First record of Elongatoolithid theropod eggshell from North America: the Asian oogenus Macroelongatoolithus from the Lower Cretaceous of Utah: Journal of Vertebrate Paleontology, v. 20, n. 1, p. 130-138.

Zelenitsky, D. K., and Hills L. V., 1996, An egg clutch of Prismatoolithus levis oosp. nov. From the Oldman Formation (Late Cretaceous), Devil’s Coulee, southern Alberta: Canadian Journal of Earth Science, v. 33, p. 1127-1131.

Zelenitsky, D. K, and Hills L. V., 1997, Normal and Pathological eggshells of Spheroolithus albertensis, oosp. Nov., from the Oldman Formation (Judith River Group, Late Cretaceous), Southern Alberta: Journal of Vertebrate Paleontology, v. 17, n. 2, p. 167-171.

Zelenitsky, D. K., Hills L. V., and Currie P. J., 1996, Parataxonomic classification of ornithoid eggshells fragments from the Oldman Formation (Judith River Group: Upper Cretaceous), southern Alberta: Canadian Journal of Earth Science, v. 33, p. 1655-1667.

Zeng, D. M., and Zhang, J. J., 1079, Fossil dinosaur eggs from the Western part of the Dongting basin, Hunan Province: Vertebrata PalAsiatica, v. 17, n. 2, p. 131-137.

Zhao, D-M., and Zhang J.-J., 1979, On the dinosaurian eggs from the Western Dongting Basin, Hunan: Vertebrata PalAsiatica,, v. 17, n. 2, p. 131-136.

Zhao, H., and Zhao Z., 1998, Dinosaur eggs from Xichuan Basin, Henan Province: Vertebrata PalAsiatica, v. 36, n. 4, p. 282-296.

Zhao, H., and Zhao X.-K., 1999, A new form of Elongatoolithid dinosaur eggs from the Lower Cretaceous Shahai Formation of Heishan, Liaoning Province: Vertebrata PalAsiatica, v. 37, n. 4, p. 278-284.

Zhao, Z.-K., 1975, The microstructure of the dinosaurian eggshells of Nanshiung, Kwangtung: Vertebrata PalAsiatica,, v. 13, n. 2, p. 105-117.

Zhao, Z.-K., 1978, A preliminary investigation of the thinning of the dinosaurian eggshells of the late Cretaceous and some related problems: Vertebrata PalAsiatica, v. 16, p. 213-221.

Zhao, Z-K., 1979, Discovery of the Dinosaurian Eggs and Footprint from Neixiang County, Henan Province: Vertebrata PalAsiatica, v. 17, n. 4, p. 304-309.

Zhao, Z.-K., 1994, Dinosaur eggs in China. On the structure and evolution of eggshells: In: Dinosaur Eggs and Babies, Edited by Carpenter K., Hirsch K. F., and Horner J. R., Cambridge University Press, p. 184-203.

Zhao, Z.-K, 2000, Nesting behavior of dinosaurs as interpreted from the Chinese Cretaceous dinosaur eggs: In: 2000 International Dinosaur Symposium for Kosong County in Korea, edited by Lee Y.-N., Proceedings of the International Dinosaur Symposium, The Paleontological Society of Korea, special publication n. 4, p. 115-126.

Zhao, Z-K, and Ding S., 1976, Discovery of the dinosaurian eggshells from Alxa, Ningxia and its stratigraphic significance: Vertebrata PalAsiatica, v. 14, p. 42-44.

Zhao, Z-K., and Jiang Y. K., 1974, Microscopic studies on the dinosaurian eggshells from Laiyang, Shandong Province: Scientia Sincia, v. 17, p. 73-83.

Zhao, Z-K., and Li. Z., 1988, A new structural type of the Dinosaur eggs from Anlu County, Hubei Province: Vertebrata PalAsiatica,, v. 26, n. 2, p. 107-115.

Zhao, Z.-K., and Li R., 1993, First record of Late Cretaceous Hypsilophodontid eggs from Bayan Manduhu, Inner Mongolia: Vertebrata PalAsatica, v. 31, n. 2, p. 77-84.

Zhao, Z., Ye, J., LI H., Zhao Z., and Zhang Y., 1991, Extinction of the Dinosaurs across the Cretaceous-Tertiary boundary in Nanxiong Basin, Guangdong Province: Vertebrata PalAsiatica, v. 29, n. 1, p. 28-29.

Zhao, Z., Ye J., Li H., Zhao Z., and Yan Z., 1991, Extinctions of the dinosaurs across the Cretaceous-Tertiary boundary in Nanxiong Basin, Guangdong Province: Vertebrata PalAsiatica, v. 29, p. 1-20.