Genus: Beelzebufo EVANS, JONES & KRAUSE, 2008
Etymology: Greek, Beel'zebul, "devil" and Latin, bufo, "toad": in reference to the size and probable life appearance of the anurans.

Species: ampinga EVANS, JONES & KRAUSE, 2008
Etymology: Malagasay, ampinga, "shield," in reference to cranial hyperossification.

Holotype: UA 9600

Locality: MAD93-25, 15°54'17.714"S, 46°34'55.946"E, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material: Fused cervical (= atlantal), and second parasacral centra.

Referred material:

UA 1959: Right squamosal, with maxillary process.

UA 2003: Sacral vertebrae.

UA 2510: Left maxilla.

UA 2513: Left frontoparietal, anterior region.

UA 2536: Right squamosal,otic process.

UA 9622: Left premaxilla.

UA 9625: Immature right nasal, maxillary process.

UA 9628: Left tibiofibula.

UA 9629: Partial left nasal.

UA 9636: Urostyle.

UA 9639: Left squamosal, partial maxillary process.

UA 9674: Right nasal, rostral proceses.

UA 9675: Left frontoparietal and exoocipital.

EVANS, GROENKE, JONES, TURNER & KRAUSE, 2014

UA 9945: Maxilla fragment.

UA 9946: Fragment of right squamosal, including suture with frontoparietal.

 

Locality: MAD93-01, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9614: posteroventral process of right squamosal.

UA 9615: Cranial fragment from antorbital margin, either nasal or frontoparietal.

FMNH PR 2003: Right half of sacral vertebra.

 

Locality: MAD93-06, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9618: Fragment of left quadratojugal.

UA 9619: Vertebral spine table.

 

Locality: MAD93-14, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9620: Fragment of ?dorsl bony plate.

 

Locality: MAD93-17, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2498: Cranial fragment.

FMNH PR 2536: Right fused squamosal-quadratojugal flange.

FMNH PR 2537: Squamosal fragment.

 

Locality: MAD93-33, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2497: Craial or vertebral spine table fragment.

UA 8677: Partial right angulosplenial.

 

Locality: MAD93-34, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9631: Fragment of right squamosal, including suture with frontoparietal.

UA 9632: Cranial ro vertebral spine table fragment.

FMNH PR 2499: Partial right maxilla.

FMNH PR 2500: Partial posterior process of right quadratojugal.

FMNH PR 2501: partial right quadratojugal.

 

Locality: MAD93-35, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9623: Fragment of otic and ventral process of squamosal.

UA 9624: Posterior process of left quadratojugal.

UA 9625: Posteriorvenral process of right squamosal.

UA 9626: Cranial fragment.

UA 9627: Partial vertebral spine taple.

UA 9635: Anterior fragment of right maxilla.

UA 9676: Fragment of right maxilla bearing maxillary nerve canal.

UA 9677: Fragment of frontoparietal or squamosal.

UA 9679: Cranial fragment.

UA 9947: Presacral vertebra.

FMNH PR 1960: Partial right premaxilla.

FMNH PR 2504: Vertebral centrum and partial neural arch.

FMNH PR 2505: Facial process of right maxilla.

FMNH PR 2506: Partial right maxilla.

FMNH PR 2507: Facial process and pars dentalis of right maxilla.

FMNH PR 2508: Cranial fragment from antorbital margin, frontoparietal or nasal.

FMNH PR 2509: Conjoined midline frontoparietal fragment.

FMNH PR 2510: Partial left maxilla.

 

Locality: MAD93-36, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9678: Vertebral spine table.

UA 9949: Fragment of otic plate of right squamosal.

 

Locality: MAD93-37, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 1959: Partial right quadratojual.

UA 52: UA 9950: Fragment of otic and ventral porcess of squamosal.

UA 9951: Left nasal fragment.

 

Locality: MAD93-73, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9622: Partial left premaxilla.

 

Locality: MAD96-21, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9628: Right tibiofibula.

 

Locality: MAD96-24, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9629: 4 fragments of large left squamosal.

 

Locality: MAD98-25, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

FMNH PR 2512: Fragmentary skull and axial column; braincase, partial frontoparietal, and right posterior skull, portionsof the left posterior skull and rostrum, left pars facialis of maxilla, partial stapes, atlas and second presacral vertebra, presacral vertebra interpreted as PS4, presacral spinosu process interpreted as PS 5 partial presacral vertebral centrum, and partial anterior urogystal.

 

Locality: MAD99-14, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9633: Right frontoparietal or nasal.

 

Locality: MAD99-29, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9634: Partial right maxilla.

 

Locality: MAD01-15, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9952: Cranial or vertebral spine table fragmetn.

 

Locality: MAD03-05, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9636: Partial urostyle.

UA 9637: Cranial ro vertebral spine table fragments.

 

Locality: MAD03-10, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9638: Otic plate of right squamosal.

 

Locality: MAD03-18, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9617: Posteriormost tip of left quadratojugal.

UA 9640: Fragment of left frontoparietal and occipital.

 

Locality: MAD05-28, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9539: Midportion of left quadratojugal.

 

Locality: MAD05-64, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9675: Partial left frontoparietal and otoccipital.

 

Locality: MAD07-15, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9674: Posterior process of left quadratojugal.

 

Locality: MAD07-20, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9953: Partial maxilla.

 

Locality: MAD10-13, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9954: Presacral vertebral column.

UA 9955: 2 cranial fragments.

 

Locality: MAD10-24, Berivotra field study area, Mahanjanga Province, Madagascar.

Horizon: Anembalemba Member, Maevarano Formation.

Biostratigraphy:

Age: Maastrichtian Stage, Senonian Subepoch, Upper Gulf Epoch, Late Cretaceous.

Material:

UA 9957: Right tibiale-fibular.

US 9958: Left nasal fragment.