EUROPEAN RUSSIA
?KOSTROMA REGION
NEISKII DISTRICT
MAKAR'EV (?) LOCALITY
RYBINSKIAN HORIZON
VETLUGA SERIES
SHILIKHA FORMATION
OLENEKIAN STAGE?
UPPER SCYTHIAN EPOCH
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)
Benthosuchus bystrowi
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EASTERN EUROPEAN RUSSIA
KOSTROMA REGION (KOSTROMSKOJ OBLAST)
NEJSKIJ RAJON
UNZHA SITE NEAR MAKARYEV
VETLUGA SUBHORIZON
RYBINSKIAN HORIZON
EARLY TRIASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LABYRINTHODONTIA OWEN, 1860
SUPERORDER: TEMNOSPONDYLIA ZITTEL, 1887
MICROORDER: LIMNARCHIA YATES & WARREN, 2000
ORDER: TREMATOSAURIA PIVETEAU, 1955
Superfamily: TREMATOSAUROIDEA Save-Soderbergh, 1935
Family: BENTHOSUCHIDAE Efremov, 1931
Subfamily: BENTHOSUCHINAE Efremov, 1931 (sensu Efremov, 1937)