SPAIN
MAESTRAZGO BASIN
PENAGOLOSA SUB-BASIN
TERUEL PROVINCE
ARAGON
EL CASTELLAR MUNICIPALITY
BERZAL NORTE (CT-10)
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
CLADE NAME: NEOSTEGOSAURIA SANCHEZ-FENOLLOSA & CABOS, 2025
Subfamily: DACENTRURINAE Olshevsky & Ford, 1993 emend Mateus, Maidment & Christiansen, 2009
Incertae sedis
Gen. sp indet.
________________________________________________________________________________________________________________________________
SPAIN
MAESTRAZGO BASIN
GALVE SUB-BASIN
TERUEL PROVINCE
ARAGON
MONTEGUADO DEL CASTILLO MUNICIPALITY
MC-2 SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
CLADE NAME: NEOSTEGOSAURIA SANCHEZ-FENOLLOSA & CABOS, 2025
Subfamily: DACENTRURINAE Olshevsky & Ford, 1993 emend Mateus, Maidment & Christiansen, 2009
Incertae sedis
Gen. sp indet.
________________________________________________________________________________________________________________________________
SPAIN
SOUTH-IBERIAN BASIN
TERUEL PROVINCE
ARAGON
RIODEVA MUNICIPALITY
SAN LORENZO 2 (RD -29)
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
CLADE NAME: NEOSTEGOSAURIA SANCHEZ-FENOLLOSA & CABOS, 2025
Subfamily: DACENTRURINAE Olshevsky & Ford, 1993 emend Mateus, Maidment & Christiansen, 2009
Incertae sedis
Gen. sp indet.
________________________________________________________________________________________________________________________________
SPAIN
SOUTH-IBERIAN BASIN
TERUEL PROVINCE
ARAGON
UNKNOWN SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
CLADE NAME: NEOSTEGOSAURIA SANCHEZ-FENOLLOSA & CABOS, 2025
Subfamily: DACENTRURINAE Olshevsky & Ford, 1993 emend Mateus, Maidment & Christiansen, 2009
Incertae sedis
Gen. sp indet.
________________________________________________________________________________________________________________________________
SPAIN
SOUTH-IBERIAN BASIN
TERUEL PROVINCE
ARAGON
RIODEVA MUNICIPALITY
EL VAGO DE BLASCO (RD -17)
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
CLADE NAME: NEOSTEGOSAURIA SANCHEZ-FENOLLOSA & CABOS, 2025
Subfamily: DACENTRURINAE Olshevsky & Ford, 1993 emend Mateus, Maidment & Christiansen, 2009
Incertae sedis
Gen. sp indet.
________________________________________________________________________________________________________________________________
SPAIN
SOUTH-IBERIAN BASIN
TERUEL PROVINCE
ARAGON
RIODEVA MUNICIPALITY
PINO DE JARQUE 2 SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
SOUTH-IBERIAN BASIN
TERUEL PROVINCE
ARAGON
RIODEVA MUNICIPALITY
PINO DE JARQUE 2 SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
MAESTRAZGO BASIN
PENAGOLOSA SUB-BASIN
TERUEL PROVINCE
ARAGON
MORA DE RUBEILOS MUNICIPALITY
EL BATAN
MR-8 FOSSIL SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
SOUTHWESERN IBERIAN RANGE
TERUEL PROVINCE
VEGUILLAS DE LA SIERRA
VS-1 SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: DRYOMORPHA SERNEO, 1986
Family: DRYOSAURIDAE Milner & Norman, 1984
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
SOUTHWESERN IBERIAN RANGE
TERUEL PROVINCE
VEGUILLAS DE LA SIERRA
VS-1 SITE
VILLAR DEL ARZOBISPO FORMATION
LATE KIMMERIDGIAN-TITHONIAN STAGE
MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
TERUEL PROVINCE
ARAGON
RIODEVA MUNICIPALITY
ESTAN DE COLON (RD-34)
VILLAR DEL ARZOBISPO FORMATION
KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
Subfamily: KENTROSAURINAE nova
Dacentrurus armatus
_____________________________________________________________________________________
SPAIN
TERUEL PROVINCE
LA PUEBAL DE VALVERDE MUNICIPALITY
VILLAR DEL ARZOBISPO FORMATION
KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
ICHNOSUPERORDER: NEOTHEROPODOPEDIDA NOVA
ICHNOORDER: NEOTETANURIPEDIDII NOVA
ICHNOSUBORDER: AVETHEROPODIPIDA NOVA
ICHNOMICRORDER: COELUROSAURIPIDIDA NOVA
Ichnofamily: GRALLATORIDAE emend Lull, 1904
Kayentapus sp
ICHNOCOHORT: ORNITHISCHIFORMEIPIDA NOVA
ICHNOMAGNORDER: ORNITHISCHIPIDA NOVA
ICHNOSUPERORDER: NEORNITHISCHIPIDA NOVA
ICHNOGRANDORDER: ORNITHOPODIPDIA NOVA
ICHNOMICRORDER: EUORNITHOPODIPIDA NOVA
ICHNOORDER: IGUANODONTIPIDIA NOVA
ICHNOSUBORDER: ANKYLOPOLLEXIPEDIDA NOVA
ICHNOINFRAORDER: STYRACOSTERNIPIDA NOVA
ICHNOPARVORDER: HADROSAURIFORMIPEDIDA NOVA
Ichnosuperfamily: IGUANODONTIDEA Nova
Ichnofamily: IGUANODONTIPODIDAE emend Vylov, 1988
Gen. sp indet.
_____________________________________________________________________________________
SPAIN
TERUEL PROVINCE
ARAGON
EL CASTELLAR MUNICIPALITY
SAN CRISTOBAL
VILLAR DEL ARZOBISPO FORMATION
KIMMERIDGIAN-TITHONIAN STAGE
UPPER MALM EPOCH
LATE JURASSIC
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: STEGOSAUROMORPHA COOPER, 1985 (EMEND)
ORDER: STEGOSAURIA MARSH, 1877
Family: STEGOSAURIDAE Marsh, 1877
Subfamily: KENTROSAURINAE nova
Dacentrurus armatus
_____________________________________________________________________________________