ENGLAND

KENT COUNTY

FOLKESTONE

GAULT

CAMBRIDGE GREENSAND

CENOMANIAN STAGE

UPPERMIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

Incertae sedis

Gen. sp indet.

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)

Family: ORNITHOCHEIRIDAE Seeley, 1870

Incertae sedis

*Ornithocheirus daviesii
= Pterodactylus daviesii
= Lonchodectes daviesii
= Ornithochirus daviesii
Ornithocheirus sp

ENGLAND

KENT COUNTY

LOWER CHALK

CENOMANIAN STAGE

UPPER MIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

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ENGLAND

KENT COUNTY

CHALK MARL

CENOMANIAN STAGE

UPPER MIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

Incertae sedis

non Polyptychodon, P. continus
= Polyptychodon mackesoni
Polyptychodon sp

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ENGLAND

KENT COUNTY

UPPER CHALK

CENOMANIAN STAGE

MIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988

Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988

Family: CHELONIIDAE Oppel, 1811

Subfamily: CHELONINAE Oppel, 1811

Chelone camperi

LEPIDOSAUROMORPHA: BENTON, 1985

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

ENGLAND

KENT COUNTY

PLENUS MARLS

CENOMANIAN STAGE

UPPER MIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

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ENGLAND

KENT COUNTY

BURHAM

CULAND PITS

LOWER CHALK-UPPER CHALK

CENOMANIAN-TURONIAN STAGE

UPPER GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE-LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

Incertae sedis

Genus sp indet.

COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988

MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988

SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987

GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988

MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988

ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988

SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988

INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988

Superfamily: CHELONIOIDEA Oppel, 1811

Family: CHELONIIDAE Oppel, 1811

Incertae sedis

Gen sp indet.

Subfamily: CHELONINAE Oppel, 1811

Chelone camperi
Chelone sp
= Chelone/Lytolma sp

Superfamily: DERMOCHELYOIDEA Willams, 1950

Family: PROTOSTEGIDAE Cope, 1889

Subfamily: CHELOSPHARGINAE Zangerl, 1953

*Cimochelys benstedi

Subfamily: PROTOSTEGINAE Cope, 1889

Tribe: PROTOSTEGINI Cope, 1889 (sensu Hooks, 1998)

Subtribe: PROTOSTEGINA Cope, 1889 (sensu Hooks, 1998)

Protostega sp

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: EURYAPSIDA COLBERT, 1945

SUPERORDER: SAUROPTERYGIA OWEN, 1860

GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994

ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835

INFRAORDER: PLESIOSAUROMORPHA NOVA

EUPLESIOSAURIA O’KEEFE, 2001

Superfamily: CRYPTOCLEIDOIDEA Williston, 1925

TRICLEDIA O’KEEFE , 2001

Family: CIMOLIASAURIDAE Delair, 1959

Cimoliasaurus smithi

INFRAORDER: PLIOSAUROMORPHA NOVA

Superfamily: PLIOSAUROIDEA Seeley, 1874

Family: PLIOSAURIDAE Seeley, 1874

Polyptychodon interruptus

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Superfamily: PTERANODONTIDEA Marsh, 1876

Incertae sedis

*Cimoliopterus cuvieri = Anhanguera cuvieri
= Pterodactylus cuvieri
= Coloborhynchus cuvieri
= Ornithochirus cuvieri
= Ptenodactylus cuvieri

Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)

Family: ORNITHOCHEIRIDAE Seeley, 1870

Incertae sedis

non Ornithocheirus, *O. giganteus
= Pterodactylus giganteus
= Lonchodectes giganteus
= Ornithochirus giganetus
= Ornithodesmus giganteus

 

_____________________________________________________________________________________

ENGLAND

KENT COUNTY

BURHAM

CULAND PITS

ZIG-ZAG CHALK FORMATION [SOUTH DOWNS NOMENCLATURE] (LOWER CHALK, ABBOTTS CLIFF CHALK FORMATION [NORTH DOWNS NOMENCLATURE]), LOWER CHALK-UPPER CHALK.

CALYCOCERAS GUERANGERI ZONE

UPPER CENOMANIAN STAGE

UPPERMOST GALLIC SUBEPOCH

LOWER GULF EPOCH

LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: DOLICHOSAURIDAE Gervais, 1852

Tribe: DOLICHOSAURINI Gervais, 1852

*Dolichosaurus longicollis

Infrafamily: NATANIA

Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)

Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)

Mosasaurus gracilis

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ENGLAND

KENT COUNTY

TOWARDS FOLKESTONE

END OF THE ROUND DOWN TUNNEL

GREY LOWER CHALK

CENOMANIAN STAGE

GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

NEODIAPSIDA: INCERTAE SEDIS

OR AMNIOTA: INCERTAE SEDIS

MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova

SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999

GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835

MICRORDER: LONGIPINNATI HUENE, 1948

ORDER: MERRIAMOSAURIA MONTANI, 1999

SUBORDER: EUICHTHYOSAURIA MONTANI, 1999

INFRAORDER: PARVIPELVIA MONTANI, 1999

PARVORDER: NEOICHTHYOSAURIA SANDER, 2000

SUPERFAMILY: OPHTHALMOSAURIDEA SEELEY, 1874

Family: OPTHALMOSAURIDAE Seeley, 1874    

Subfamily: PLATYPTERYGIINAE Arkhangelsky 2011

Pervushovisaurus campylodon = Platypterygius campylodon
= Ichthyosaurus campylodon
= Myopterygius campylodon

 

 

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ENGLAND

KENT COUNTY

BURHAM

WOULDHAM

PETER’S PIT

LOWER CHALK

HOLBASTER SUBGLOBOSUS ZONE

CENOMANIAN STAGE

UPPER GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: DOLICHOSAURIDAE Gervais, 1852

Tribe: DOLICHOSAURINI Gervais, 1852

Coniasaurus crassidens

_____________________________________________________________________________________

ENGLAND

KENT COUNTY

CHARING

HART HILL

CENOMANIAN STAGE

UPPER GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

INFRACLASS: NEODIAPSIDA BENTON, 1985

LEGION: LEPIDOSAUROMORPHA BENTON, 1983

SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)

MAGNORDER: SQUAMATA, OPPEL, 1811

SUPERORDER: LACERTILIA OWEN, 1842

GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961

MICRORDER: SCINCOMORPHA CAMP, 1923

ORDER: ANGUIMORPHA FURBRINGER, 1900

SUBORDER: DIPLOGLOSSA COPE, 1864

INFRAORDER: THECAGLOSSA COPE, 1890

PARVORDER: MONSTERSAURIA NORELL & GAO, 1977

Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)

Family: MOSASAURIDAE Gervais, 1853

Subfamily: DOLICHOSAURIDAE Gervais, 1852

Tribe: DOLICHOSAURINI Gervais, 1852

Coniasaurus crassidens

_____________________________________________________________________________________

ENGLAND

KENT COUNTY

WOULDHAM

PETER’S QUARRIES

LOWER PART OF THE GREY CHALK FORMATION

CENOMANIAN-TURONIAN STAGE

UPPER GALLIC SUBEPOCH

LOWER GULF EPOCH

MIDDLE-LATE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: CHELONOMORPHA KUHN, 1960

SUPERCOHORT: TESTUDINES LINNAEUS, 1758

Incertae sedis

Genus sp indet.

_____________________________________________________________________________________

ENGLAND

KENT COUNTY

WOULDHAM

PETER’S PIT

GREY CHALK FORMATION

HOLASTER SUBGLOBOSUS ZONE

CENOMANIAN STAGE

UPPER MIDDLE GALLIC SUBEPOCH

LOWERMOST GULF EPOCH

MIDDLE CRETACEOUS

STEM: TETRAPODA GOODRICH, 1930

SUPERCLASS: AMNIOTA HAECKEL, 1866

CLASS: REPTILIA LAURENTI, 1768

SUBCLASS: DIAPSIDA OSBORN, 1904

SUPERLEGION: NEODIAPSIDA BENTON, 1985

LEGION: ARCHOSAUROMORPHA HUENE, 1946

SUPERCOHORT: AVEMETATARSALIA BENTON, 1999

COHORT: PROLACERTIFORMES CAMP, 1945 NOVA

SUPERORDER: TAPINOPLATIA PETERS, 2000

GRANDORDER: CHARACIOPODA PETERS, 2000

MICRORDER: FENESTRASAURIA PETERS, 2000

ORDER: PTEROSAURIA KAUP, 1834

SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994

Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)

Family: ORNITHOCHEIRIDAE Seeley, 1870

Incertae sedis

Ornithocheirus sp

_____________________________________________________________________________________