MARYLAND
PRINCE GEORGE’S COUNTY
LANDOVER
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904 OSBORN, 1903
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: CIMOLIASAURIDAE Delair, 1959
Cimoliasaurus sp
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: HALISAURINAE Bardet, Pereda Suberbiola, Iarochene, Bouya & Amaghzaz, 2005
Halisaurus platyspondylus
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Family: GAVIALIDAE Cuvier, 1807
Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)
Thoracosaurus neocesariensis
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
N OF KERBY HILL FARM ON INDIAN HEAD HIGHWAY (MD-210)
S OF OXON HILL
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Subfamily: HALISAURINAE Bardet, Pereda Suberbiola, Iarochene, Bouya & Amaghzaz, 2005
Halisaurus platyspondylus
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
HAMPTON MALL IN PRINCE
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988
Family: OSTEOPYGIDAE Zangerl, 1953
Osteopygis emarginatus
= Lytoloma angusta?
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOC.A
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus hoffmanni
= Mosasaurus maximusLEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus? antiquus
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOCALITY B
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988
Family: OSTEOPYGIDAE Zangerl, 1953
Osteopygis emarginatus
= Lytoloma angustaSuperfamily: DERMOCHELYOIDEA Willams, 1950
Family: TOXOCHELYIDAE Baur, 1895
Subfamily: LOPHOCHELYINAE Zangerl, 1953
Peritresius ornatus
= Chelone ornataINFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Trionyx priscis
= Amyda priscaSUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: CIMOLIASAURIDAE Delair, 1959
Cimoliasaurus magnus
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus hoffmanni
= Mosasaurus maximus
Mosasaurus lemonnieri
= Mosasaurus conodon
= Clidastes conodonLEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Family: GAVIALIDAE Cuvier, 1807
Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)
Thoracosaurus neocesariensis
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOCALITY C
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: EUCHELONIOIDEA Gaffney & Meylan, 1988
Family: OSTEOPYGIDAE Zangerl, 1953
Osteopygis emarginatus
= Lytoloma angusta ?Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: TOXOCHELYIDAE Baur, 1895
Subfamily: LOPHOCHELYINAE Zangerl, 1953
Peritresius ornatus
= Chelone ornataINFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Trionyx priscis
= Amyda priscaSUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus hoffmanni
= Mosasaurus maximus
Mosasaurus dekayiLEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Family: GAVIALIDAE Cuvier, 1807
Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)
Thoracosaurus neocesariensis
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus? antiquus
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOC D
SERVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Family: GAVIALIDAE Cuvier, 1807
Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)
Thoracosaurus neocesariensis
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOCALITY E
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: TOXOCHELYIDAE Baur, 1895
Subfamily: LOPHOCHELYINAE Zangerl, 1953
Peritresius ornatus
= Chelone ornateSUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Ornithomimus? antiquus
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOCALITY F
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: TOXOCHELYIDAE Baur, 1895
Subfamily: LOPHOCHELYINAE Zangerl, 1953
Peritresius ornatus
= Chelone ornate
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
LOCALITY G
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: TOXOCHELYIDAE Baur, 1895
Subfamily: LOPHOCHELYINAE Zangerl, 1953
Peritresius ornatus
= Chelone ornateSUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MARYLAND
PRINCE GEORGE’S COUNTY
NEAR THE TOWN OF BOWIE
SEVERN FORMATION
EARLY LATE MAASTRICHTIAN STAGE
UPPERMOST SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
Gen. sp indet.
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: TOXOCHELYIDAE Baur, 1895
Subfamily: LOPHOCHELYINAE Zangerl, 1953
Peritresius ornatus
= Chelone ornataINFRAORDER: CHELOMACRYPTODIRA GAFFNEY & MEYLAN, 1988
PARVORDER: TRIONYCHIDA GRAY, 1870
Superfamily: TRIONYCHIDEA Bell, 1828 (sensu Gray, 1870)
Family: TRIONYCHIDAE Bell, 1828
Trionyx priscis
= Amyda priscaSUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: EURYAPSIDA COLBERT, 1945
SUPERORDER: SAUROPTERYGIA OWEN, 1860
GRANDORDER: EOSAUROPTERYGIA emend RIEPPEL, 1994
ORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
SUBORDER: PLESIOSAURIA OWEN, 1840 NON BLAINVILLE, 1835
INFRAORDER: PLESIOSAUROMORPHA NOVA
EUPLESIOSAURIA O’KEEFE, 2001
Superfamily: CRYPTOCLEIDOIDEA Williston, 1925
TRICLEDIA O’KEEFE , 2001
Family: CIMOLIASAURIDAE Delair, 1959
Cimoliasaurus magnus
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: ANGUIMORPHA FURBRINGER, 1900
SUBORDER: DIPLOGLOSSA COPE, 1864
INFRAORDER: THECAGLOSSA COPE, 1890
PARVORDER: MONSTERSAURIA NORELL & GAO, 1977
Superfamily: VARANOIDEA Gray, 1827 (sensu Camp, 1923)
Family: MOSASAURIDAE Gervais, 1853
Infrafamily: NATANIA
Subfamily: MOSASAURINAE Gervais, 1853 (sensu Williston, 1897)
Tribe: MOSASAURINI Gervais, 1853 (sensu Russell, 1967)
Mosasaurus
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: EUSUCHIA emend HUXLEY, 1875
INFRAORDER: CROCODYLIA emend GMELIN, 1877
Family: GAVIALIDAE Cuvier, 1807
Subfamily: THORACOSAURINAE Cope, 1875 (sensu Nopcsa, 1928)
Thoracosaurus neocesariensis
PARVORDER: BREVIROSTRES
Superfamily: ALLIGATORIDEA Buffetaut, 1985
GLOBIDONTA BROUCHU, 1999
Family: ALLIGATORIDAE Gray, 1844
Bottosaurus harlani
= Crocodilus harlaniDeinosuchus rugosus
Subfamily: ALLIGATORINAE Gray, 1844 (sensu Kalin, 1940)
Tribe: ALLIGATORINI nova
Allognathosuchus sp
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
SUBORDER: ORNITHOMIMOSAURIA BARSBOLD, 1976
Family: ORNITHOMIMIDAE Marsh, 1890
Incertae sedis
Gen. sp indet.
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
INFRAORDER: STYRACOSTERNA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
SUPERFAMILY: HADROSAUROIDEA COPE, 1869 (SENSU SERENO, 1986)
Family: HADROSAURIDAE COPE, 1869
Incertae sedis
Gen. sp indet.
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