MADAGASCAR
MAHAJANGA
MAJUNGA PROVINCE
BERIVOTRO
BERIVOTRO FORMATION
SENONIAN OR CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
*Majungasaurus crenatissimus
= Megalosaurus crenatissimusCOHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
Incertae sedis
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: ZATHERIA MCKENNA, 1975
INFRALEGION: INCERTAE SEDIS
Genus: Nova
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
JUSE EAST OF BERIVOTRA VILLAGE
MAEVARENO SANDSTONE
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: INCERTAE SEDIS
Gen. sp indet.
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
Incertae sedis
Gen. sp indet.
SUBORDER: GONDWANATHERIA Mones, 1987
Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984
Lavanify miolaka
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR THE VILLAGE OF BERIVOTRA
MAD96-01
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: THERIIFORMES (ROWE, 1988) McKENNA & BELL, 1997
INFRACLASS: ALLOTHERIA (MARSH, 1880) McKENNA & BELL, 1997
ORDER: MULTITUBERCULATA COPE, 1884
SUBORDER: GONDWANATHERIA Mones, 1987
Family: SUDAMERICICIDAE Scillato-Yane & Prasad, 1984
*Lavanify miolaka
_____________________________________________________________________________________
N. W. MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA BENTON, 1983
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA, OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
GRANDORDER: SCINCOGEKKONOMORPHA SUKHANOV, 1961
MICRORDER: SCINCOMORPHA CAMP, 1923
ORDER: LEPTOGLOSSA COPE, 1900
SUBORDER: ZONUROMORPHA ALIFANOV, 2000
INFRAORDER: ZONURIA FURBRINGER, 1900
Superfamily: CORDYLOIDEA Fritzinger, 1826
Family: CORDYLIDAE? Gray, 1837
Gen. sp indet.
_____________________________________________________________________________________
NORTH-WEST MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
APPROXIMATELY 1 KM SOUTH-EAST OF THE VILLAGE OF BERIVOTRA
MAEVARANO FORMATION
NEAR THE TOP OF THE CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
Also a long snouted form.
Incertae sedis
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: PEIROSAURIDAE Gasparini, 1982
Gen. nova?
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
*Mahajangasuchus insignis
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
*Miadanasuchus oblita
= Trematochampsa oblita
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MASIAKAKOHO STUDY AREA
MAD 05-61
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
Incertae sedis
Sokatra antitra
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
BERIVOTRA STUDY AREA
MAD 07-66
MAEVARANO FORMATION
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
Incertae sedis
Sokatra antitra
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-01
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
NEAR THE VILLAGE OF BERIVOTRA
MAD93-18
MAEVARANO FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: TETANURAE GAUTHIER, 1986
MICRORDER: COELUROSAURIA HUENE, 1914
ORDER: MANIRAPTORFORMES HOLTZ, 1996
AVETHEROPOA PAUL, 2002
AVEPECTORA PAUL, 2002
SUBORDER: MANIRAPTORA GAUTHIER, 1986
INFRAORDER: PARVES SERENO, 1997
PARVORDER: DEINONYCHOSAURIA COLBERT & D. A. RUSSELL, 1969
Family: DROMAEOSAURIDAE Russell, 1969
Subfamily: UNENLAGIINAE Bonaparte, 1999
*Rahonavis ostromi
= Rahona ostromi
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-33
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 93-34
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PODOCNEMIDERA Cope, 1868
Incertae sedis
Sokatra antitra
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-35
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Hyperfamily: PELOMEDUSERA Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Subfamily: Erymnochelus group
Cf. Erymnochelys
Hyperfamily: PODOCNEMIDERA Cope, 1868
Incertae sedis
*Sokatra antitra
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
*Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-79
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Bothremyididae gen sp indet
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-37
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: AMPHIBIA LINNAEUS, 1758
SUBCLASS: LISSAMPHIBIA, HAECKEL, 1866?
SUPERORDER: SALIENTIA LAURENTI, 1768
ORDER: ANURA RAFINESQUE, 1815
SUBORDER: MESOBATRACHIA LAURENT, 1979
Superfamily: PELOBATOIDEA Stanius, 1856 , nom. corr. Spinar, 1971 (sensu Bolkay, 1919)
Family: PELOBATIDAE Stanius, 1856 , nom. corr. Spinar, 1971
Gen. sp indet.
_____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
MAHAJANGA BASIN
NEAR BERIVOTRA
MAD 93-79
MAEVARANO FORMATION
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
UPPER GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGNORDER: PLEURODIRA GAFFNEY & MEYLAN, 1988
ORDER: PLEURODIRIDA COPE, 1864
SUBORDER: EUPLEURODIRA GAFFNEY & MEYLAN, 1988
Superfamily: PELOMEDUSOIDES Cope, 1868 Cope, 1868
Family: PELOMEDUSIDAE Cope, 1868
Subfamily: PELOMEDUSINAE Cope, 1868
Bothremyididae gen sp indet
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 95-07
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAHAJANGA PROVINCE
WHICH LIE APPROXIMATELY 4-5 KM SOUTHEAST OF THE MIAN FIELD STUDY AREA SURROUNDING THE VILLAGE OF BERIVOTRA
BETWEEN THE TWO MIADANA HILLS
MAD98-19
MAEVARANO FORMATION
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
ORDER: MESOSUCHIA EMEND HUXLEY, 1875
PARVORDER: METASUCHIA BENTON & CLARK, 1988
INFRAORDER: METAMESOSUCHIA HULKE, 1878
NEOSUCHIA CLARK, IN BENTON & CLARK, 1988
Family: TREMATOCHAMPSIDAE Buffetaut, 1974
Miadanasuchus oblita
= Trematochampsa oblita
_____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 99-35
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
____________________________________________________________________________________
MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
MAD 01-04
MAEVARANO FORMATION
ANEMBALEMBA MEMBER
MAASTRICHTIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: CRUROTARSI SERENO & ARCUCCI, 1990
COHORT: CROCODYLOTARSI BENTON & CLARK, 1988
MAGNORDER: SUCHIA KREBS, 1974
SUPERORDER: CROCODYLOMORPHA WALKER, 1968
GRANDORDER: CROCODILIFORMES HAY, 1930
MICRORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
SUBORDER: MESOEUROCROCODYLIA WHETSTONE & WHYBROW, 1983
MESOEUROCRODYLIA: NEW FAMILES?
GONDWANASUCHIA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIMORPHA DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
PEIROSAURIFORMES DE SOUZA CARVALHO, RIBEIRO & DOS SANTOS AVILLA, 2004
Superfamily: PEIROSAURIDAE Gasparini, 1982 emend de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Family: PEIROSAURIDAE Gasparini, 1982
Subfamily: LOMASUCHINAE de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Tribe MAHAJANGASUCHINI de Souza Carvalho, Ribeiro & dos Santos Avilla, 2004
Mahajangasuchus insignis
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MADAGASCAR
MAJUNGA PROVINCE
46 KM S OF MAJUNGA
BETSIBOKA
“GRES DE MAEVARENO”
CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Superfamily: TITANOSAUROIDEA Lydekker, 1885 (sensu Upchurch, 1999)
Family: TITANOSAURIDAE Lydekker, 1885
Incertae sedis
non Titanosaurus, T? madagascariensis
= Laplatasaurus madagascariensis
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MADAGASCAR
MAJUNGA PROVINCE
‘GRES DE MAEVANO’
PROBABLY CAMPANIAN STAGE
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
*Majungatholus atopus
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MADAGASCAR
MAJUNGA PROVINCE
MAHAJANGA BASIN
CAMPANIAN STAGE?
SENONIAN SUBEPOCH
GULF EPOCH
LATE CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1903
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: THEROPODA EMEND MARSH, 1881
AVEPODA PAUL, 2002
SUPERORDER: NEOTHEROPODA BAKKER, 1986
GRANDORDER: CERATOSAURIA NOVAS, 1992
AVEROSTRA PAUL, 2001
ORDER: CERATOSAURIA MARSH, 1884
Superfamily: ABELISAUROIDEA Novas, 1989
Family: ABELISAURIDAE Bonaparte & Novas, 1985
Subfamily: CARNOTAURINAE Sereno, 1998
Majungasaurus crenatissimus
= Megalosaurus crenatissimus
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