SOUTHERN UNITED KINGDOM
ENGLAND
KENT COUNTY
FOLKESTONE WARREN
LOWER GREENSAND
MAMMILATUM
EARLY CRETACEOUS
ICHNOSTEM: TETRAPODIPIDA NOVA
ICHNOSUPERCLASS: AMNIOTAIPIDII NOVA
ICHNOCLASS: REPTILIPEDIA VIALOV, 1966
ICHNOSUBCLASS: DIAPSIDIPIDII NOVA
ICHNOSTEM-GROUP: NEODIAPSIDIPEDII NOVA
ICHNOINFRACLASS: NEODIAPSIDIPEDII NOVA
ICHNOSUBDIVISION: ARCHOSAURIPEDIDII NOVA
ICHNOSUPERCOHORT: AVEMETATARSALIPEDII NOVA
ICHNOLEGION: DINOSAUROMORPHIPEDA NOVA
ICHNOSUBLEGION: DINOSAURIFORMEIPEDIDII NOVA
ICHNOSUPERCOHORT: DINOSAUROPEDIDA NOVA
ICHNOCOHORT: THEROPODIPEDIA NOVA
Incertae sedis
Gen. sp indet.
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ENGLAND
KENT COUNTY
WEALDEN FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
SUPERORDER: SAUROPODOMORPHA HUENE, 1932
GRANDORDER: BRONTOSAURIA OLSHEVSKY, 1991 (emend)
MICRORDER: SAUROPODA MARSH, 1878
ORDER: EUSAUROPODA
SUBORDER: NEOSAUROPODA BONAPARTE, 1986
INFRAORDER: MACRONARIA WILSON & SERENO, 1998
PARVORDER: TITANOSAURIFORMES SALGADO ET AL, 1997
Family: BRACHIOSAURIDAE Riggs, 1904
Subfamily: BRACHIOSAURINAE Riggs, 1904 (sensu Janensch, 1929)
Pelorosaurus conybearei
Pelorosaurus sp
= Ornithopsis sp
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ENGLAND
KENT COUNTY
AN ABANDONED QUARRY
MEMBER OF THE HASTINGS BEDS
NEOCOMIAN SUBEPOCH
LOWER EARLY CRETACEOUS EPOCH
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
_____________________________________________________________________________________SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: MAMMALIA LINNAEUS, 1758
SUBCLASS: PROTOTHERIA GILL, 1872
INFRACLASS: TRECHNOTHERIA McKENNA, 1975
LEGION: CLADOTHERIA McKENNA, 1975
SUBLEGION: DRYOLESTOIDEA BUTLER, 1939
ORDER: DRYOLESTOIDA PROTHERO, 1981
Family: DRYOLESTIDAE Marsh, 1879
Laolestes hodsoni
= *Melanodon hodsoni
ENGLAND
KENT COUNTY
NEAR FOLKESTONE
CHALK MARL
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
SUPERORDER: EURYOPDA SERENO, 1986
GRANDORDER: ARMATOSAURIA ZHAO, 1983 (emend)
ORDER: ANKYLOSAURIA OSBORN, 1923
SUBORDER: SCELIDOSAURIA COOPER, 1985
INFRAORDER: ANKYLOSAURA OSBORN, 1923
Family: NODOSAURIDAE Marsh, 1890
Acanthopholis sp
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ENGLAND
KENT COUNTY
TURON
MIDDLE CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ORNITHOCHEIRIDAE Seeley, 1870
*Lonchodectes compressirostris
= Pterodactylus compressirostris
= Lonchodectes compressirostris
= Ornithocheirus compressirostrisIncertae sedis
*Ornithocheirus? diomedius
= Osteornis diomedius
= Cimoliornis diomedius
= Ornithocheirus diomedius
= Ornithochirus diomedius
= Pterodactylus diomedius
= Pterodactylus conirostris
Ornithocheirus spSuperfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Ornithocheirus, O. hlavatschi
= Cretornis hlavatschi
= Ornithochirus hlavatschi
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ENGLAND
KENT COUNTY
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Subfamily: CHELONINAE Oppel, 1811
Chelone sp
SUBCLASS: DIAPSIDA OSBORN, 1904 OSBORN, 1903
INFRACLASS: NEODIAPSIDA BENTON, 1985
NEODIAPSIDA: INCERTAE SEDIS
OR AMNIOTA: INCERTAE SEDIS
MAGANORDER: ICHTHYOPTERYGIA BAUR, 1887 emend nova
SUPERORDER: EOICHTHYOSAURIA MOTANI, 1999
GRANDORDER: ICHTHYOSAURIA EMEND BLAINVILLE, 1835
Incertae sedis
Gen. sp indet.
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ENGLAND
KENT COUNTY
BURHAM CHALKPIT
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: CHELONIOIDEA Oppel, 1811
Family: CHELONIIDAE Oppel, 1811
Subfamily: CHELONINAE Oppel, 1811
Chelone sp
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ENGLAND
KENT COUNTY
BURHAM
LOWER CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Ornithocheirus, O. hlavatschi
= Cretornis hlavatschi
= Ornithochirus hlavatschi
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ENGLAND
KENT COUNTY
SNODLAND
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: PTERANODONTIDEA Marsh, 1876
Family: PTERANODONTIDAE Marsh, 1876
Incertae sedis
non Ornithocheirus, O. hlavatschi
= Cretornis hlavatschi
= Ornithochirus hlavatschi
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ENGLAND
KENT COUNTY
WHITE CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ANHANGUERIDAE Campos & Kellner 1985
Anhanguera cuvieri
= Pterodactylus cuvieri
= Coloborhynchus cuvieri
= Ornithochirus cuvieri
= Ptenodactylus cuvieri
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ENGLAND
KENT COUNTY
HALLING
LOWER CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Superfamily: ORNITHOCHEIROIDEA Seeley, 1870 (sensu Seeley, 1891)
Family: ANHANGUERIDAE Campos & Kellner 1985
Anhanguera cuvieri
= Pterodactylus cuvieri
= Coloborhynchus cuvieri
= Ornithochirus cuvieri
= Ptenodactylus cuvieri
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ENGLAND
KENT COUNTY
BURHAM
LOWER CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
COHORT: CASICHELYDIA emend GAFFNEY & MEYLAN, 1988
MAGANORDER: CRYPTODIRA GAFFNEY & MEYLAN, 1988
SUPERORDER: SELMACRYPTODIRA GAFFNEY, HUTCHISON, JENKINS & MEEKR, 1987
GRANDORDER: DAIOCRYPTODIRA GAFFNEY & MEYLAN, 1988
MICRORDER: CENTROCRYPTODIRA GAFFNEY & MEYLAN, 1988
ORDER: POLYCRPTODIRA GAFFNEY & MEYLAN, 1988
SUBORDER: PROCOELOCRYPTODIRA GAFFNEY & MEYLAN, 1988
INFRAORDER: CHELONOIDEA GAFFNEY & MEYLAN, 1988
Superfamily: DERMOCHELYOIDEA Willams, 1950
Family: PROTOSTEGIDAE Cope, 1889
Subfamily: CHELOSPHARGINAE Zangerl, 1953
Cimochelys benstedi
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ENGLAND
KENT COUNTY
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
INFRACLASS: NEODIAPSIDA BENTON, 1985
LEGION: LEPIDOSAUROMORPHA DUMERIL & BIBRON, 1839
SUBLEGION: LEPIDOSAURIA HAECKEL, 1866 (DUMERIL & BIBRON, 1839)
MAGNORDER: SQUAMATA OPPEL, 1811
SUPERORDER: LACERTILIA OWEN, 1842
Incertae sedis
*Raphiosaurus no species given
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ENGLAND
KENT COUNTY
MAIDSTONE
KENTISH RAG
GREEN-SAND FORMATION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: CHELONOMORPHA KUHN, 1960
SUPERCOHORT: TESTUDINES LINNAEUS, 1758
Incertae sedis
*Protemys serrata
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ENGLAND
KENT COUNTY
NEAR MAIDSTONE
KENTISH RAG
LOWER GREENSAND FORMTION
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
SUPERORDER: NEORNITHISCHIA COOPER, 1985
GRANDORDER: ORNITHOPODA MARSH, 1871
MICRORDER: EUORNITHOPODA SERENO, 1986
ORDER: IGUANODONTIA DOLLO, 1888
SUBORDER: ANKYLOPOLLEXIA SERENO, 1986
PARVORDER: HADROSAURIFORMES SERENO, 1997
Superfamily: IGUANODONTIDEA Cope, 1869 (sensu Sereno, 1986)
Family: IGUANODONTIDAE Cope, 1869
Mantellodon carpenteri
= *Iguanodon mantelli
= Therosaurus mantelli
= Iguanodon mantelliIncertae sedis
Gen. sp indet.
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ENGLAND
KENT COUNTY
NEAR MAIDSTONE
CHALK
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUPERCOHORT: AVEMETATARSALIA BENTON, 1999
COHORT: PROLACERTIFORMES CAMP, 1945 NOVA
SUPERORDER: TAPINOPLATIA PETERS, 2000
GRANDORDER: CHARACIOPODA PETERS, 2000
MICRORDER: FENESTRASAURIA PETERS, 2000
ORDER: PTEROSAURIA KAUP, 1834
SUBORDER: EUPTERODACTYLOIDEA BENNETT, 1994
Incertae sedis
Gen sp indet.
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ENGLAND
KENT COUNTY
QUARRY AT SOUTHBOROUGH
HASTINGS SANDS
LOWER WEALDEN
EARLY CRETACEOUS
STEM: TETRAPODA GOODRICH, 1930
SUPERCLASS: AMNIOTA HAECKEL, 1866
CLASS: REPTILIA LAURENTI, 1768
SUBCLASS: DIAPSIDA OSBORN, 1904
SUPERLEGION: NEODIAPSIDA BENTON, 1985
LEGION: ARCHOSAUROMORPHA HUENE, 1946
SUBLEGION: DINOSAUROMORPHA BENTON & NORMAN IN BENTON, 1985
INFRALEGION: DINOSAURIFORMES NOVAS, 1992
SUPERCOHORT: DINOSAURIA OWEN, 1842
COHORT: ORNITHISCHIFORMES COOPER, 1985
MAGNORDER: ORNITHISCHIA SEELEY, 1888
Incertae sedis
*Thecospondylus horneri